出典(authority):フリー百科事典『ウィキペディア(Wikipedia)』「2015/08/17 16:25:29」(JST)
Branchial arches, or gill arches, are a series of bony "loops" present in fish, which support the gills. As gills are the primitive condition of vertebrates, all vertebrate embryos develop pharyngeal arches, though the eventual fate of these arches varies between taxa. In jawed fish, the first arch develops into the jaws, the second into the hyomandibular complex, with the posterior arches supporting gills. In amphibians and reptiles, many elements are lost including the gill arches, resulting in only the oral jaws and a hyoid apparatus remaining. In mammals and birds, the hyoid is still more simplified.
All basal vertebrates breathe with gills. The gills are carried right behind the head, bordering the posterior margins of a series of openings from the esophagus to the exterior. Each gill is supported by a cartilagenous or bony gill arch.[1] Bony fish have three pairs of arches, cartilaginous fish have five to seven pairs, and primitive jawless fish have seven. The vertebrate ancestor no doubt had more arches, as some of their chordate relatives have more than 50 pairs of gills.[2]
In amphibians and some primitive bony fishes, the larvae bear external gills, branching off from the gill arches.[3] These are reduced in adulthood, their function taken over by the gills proper in fishes and by lungs in most amphibians. Some amphibans retain the external larval gills in adulthood, the complex internal gill system as seen in fish apparently being irrevocably lost very early in the evolution of tetrapods.[4]
The branchial system is typically used for respiration and/or feeding. Many fish have modified anterior gill arches into pharyngeal jaws, often equipped with specialized pharyngeal teeth for handling particular prey items (long, sharp teeth in carnivorous moray eels compared to broad, crushing teeth in durophagous black carp). In amphibians and reptiles, the hyoid arch is modified for similar reasons. It is often used in buccal pumping and often plays a role in tongue protusion for prey capture. In species with highly specialized ballistic tongue movements such as chameleons or some plethodontid salamanders, the hyoid system is highly modified for this purpose, while it is often hypertrophied in species which use suction feeding. Species such as snakes and monitor lizards, whose tongue has evolved into a purely sensory organ, often have very reduced hyoid systems.
The primitive arrangement is 7 (possibly 8) arches, each consisting of the same series of paired (left and right) elements, in order of dorsal to ventral: Pharyngobranchial, epibranchial, ceratobranchial, hypobranchial, and basibranchial. The pharyngobranchials articulate with the neurocranium, while the left and right basibranchials connect to each other (often fusing into a single bone). When part of the hyoid system, the names of the bones are altered by replacing "branchial" with "hyal", thus "ceratobranchial" becomes "ceratohyal".
Higher vertebrates do not have gills. The gill arches form as pharyngeal arches during embryogenesis, and lay the basis of essential structures such as jaws, the thyroid gland, the larynx, the columella (corresponding to the stapes in mammals) and in mammals the malleus and incus.[2]
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リンク元 | 「鰓弓」「branchial region」「visceral arch」 |
拡張検索 | 「third and fourth branchial arch syndrome」 |
関連記事 | 「arc」「arch」「branchia」「branchial」 |
# | 鰓弓 | 神経 | 筋 | 骨格 | 動脈* |
1 | 顎弓(上顎隆起、下顎隆起) | CN V 三叉神経: 上顎枝、下顎枝(第一鰓弓の筋を支配) |
咀嚼筋(側頭筋、咬筋、内側翼突筋・外側翼突筋)、顎舌骨筋、顎二腹筋前腹、口蓋帆張筋と鼓膜張筋 | 上顎突起(顎前骨、上顎骨、頬骨、側頭骨の一部)、メッケル軟骨(下顎骨、キヌタ骨、ツチ骨、前ツチ骨靱帯、蝶下顎靱帯) | 消失し一部残存(顎動脈) |
2 | 舌骨弓 | CN VII 顔面神経 |
顔面表情筋(頬筋、耳介筋、前頭筋、広頚筋、口輪筋および眼輪筋)、顎二腹筋後腹、茎突舌骨筋、アブミ骨筋 | アブミ骨、茎状突起、茎突舌骨靱帯、舌骨小角と舌骨体の上部 | 消失し一部残存(舌骨動脈、アブミ骨動脈) |
3 | CN IX 舌咽神経 |
茎突咽頭筋 | 舌骨大角と舌骨体の下部 | 総頚動脈、内頚動脈の基部。外頚動脈が出芽 | |
4 | CN X 迷走神経 |
輪状甲状筋、口蓋帆挙筋 | 喉頭軟骨(甲状軟骨、輪状軟骨、披裂軟骨、小角軟骨、および楔状軟骨 | 大動脈弓の一部 | |
5 | 上喉頭神経(第四鰓弓支配神経) | 咽頭収縮筋 | 消失 | ||
6 | 反回神経(下喉頭神経)(第六鰓弓支配神経) | 喉頭内の筋 | 動脈管と肺動脈の基部 |
内臓・
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