Sensory system |
|
Typical sensory system: the visual system, illustrated by the classic Gray's FIG. 722– This scheme shows the flow of information from the eyes to the central connections of the optic nerves and optic tracts, to the visual cortex. Area V1 is the region of the brain which is engaged in vision. |
Latin |
organa sensuum |
A sensory system is a part of the nervous system responsible for processing sensory information. A sensory system consists of sensory receptors, neural pathways, and parts of the brain involved in sensory perception. Commonly recognized sensory systems are those for vision, hearing, somatic sensation (touch), taste and olfaction (smell). In short, senses are transducers from the physical world to the realm of the mind where we interpret the information, creating our perception of the world around us.[1]
The receptive field is the specific part of the world to which a receptor organ and receptor cells respond. For instance, the part of the world an eye can see, is its receptive field; the light that each rod or cone can see, is its receptive field.[2] Receptive fields have been identified for the visual system, auditory system and somatosensory system, so far.
Contents
- 1 Stimulus
- 2 Modality
- 2.1 V1 (vision)
- 2.2 A1 (auditory - hearing)
- 2.3 S1 (somatosensory - touch and proprioception)
- 2.4 G1 (gustatory - taste)
- 2.5 O1 (olfactory - smell)
- 3 Human sensory system
- 4 Diseases
- 5 See also
- 6 References
|
Stimulus
Sensory systems code for four aspects of a stimulus; type (modality), intensity, location, and duration. Arrival time of a sound pulse and phase differences of continuous sound are used for localization of sound sources. Certain receptors are sensitive to certain types of stimuli (for example, different mechanoreceptors respond best to different kinds of touch stimuli, like sharp or blunt objects). Receptors send impulses in certain patterns to send information about the intensity of a stimulus (for example, how loud a sound is). The location of the receptor that is stimulated gives the brain information about the location of the stimulus (for example, stimulating a mechanoreceptor in a finger will send information to the brain about that finger). The duration of the stimulus (how long it lasts) is conveyed by firing patterns of receptors. These impulses are transmitted to the brain through afferent neurons.
Modality
A stimulus modality (sensory modality) is a type of physical phenomenon that can be sensed. Examples are temperature, taste, sound, and pressure. The type of sensory receptor activated by a stimulus plays the primary role in coding the stimulus modality.
In the memory-prediction framework, Jeff Hawkins mentions a correspondence between the six layers of the cerebral cortex and the six layers of the optic tract of the visual system. The visual cortex has areas labelled V1, V2, V3, V4, V5, MT, IT, etc. Thus Area V1 mentioned below, is meant to signify only one class of cells in the brain, for which there can be many other cells which are also engaged in vision.
Hawkins lays out a scheme for the analogous modalities of the sensory system. Note that there can be many types of senses, some not mentioned here. In particular, for humans, there will be cells which can be labelled as belonging to V1, V2 A1, A2, etc.:
V1 (vision)
The human eye is the first element of a
sensory system: in this case, vision, for the visual system.
Visual Area 1, or V1, is used for vision, via the visual system to the primary visual cortex.
A1 (auditory - hearing)
Auditory Area 1, or A1, is for hearing, via the auditory system, the primary auditory cortex.
S1 (somatosensory - touch and proprioception)
Somatosensory Area 1, or S1, is for touch and proprioception in the somatosensory system. The somatosensory system feeds the Brodmann Areas 3, 1 and 2 of the primary somatosensory cortex. But there are also pathways for proprioception (via the cerebellum), and motor control (via Brodmann area 4).
G1 (gustatory - taste)
Gustatory Area 1, or G1, is used for taste. Taste does not refer to the flavor of food, because the flavor of the food refers to the odor. The five qualities of taste include sourness, bitterness, sweetness, saltiness, and a recently discovered protein taste quality called umami.
O1 (olfactory - smell)
Olfactory Area 1, or O1, is used for smell. In contrast to vision and hearing, the olfactory bulbs are not cross-hemispheric; the right bulb connects to the right hemisphere and the left bulb connects to the left hemisphere.
Human sensory system
The Human sensory system consists of the following sub-systems:
- Visual system consists of the photoreceptor cells, optic nerve, and V1.
- Auditory system
- Somatosensory system consists of the receptors, transmitters (pathways) leading to S1, and S1 that experiences the sensations labelled as touch or pressure, temperature (warm or cold), pain (including itch and tickle), and the sensations of muscle movement and joint position including posture, movement, and facial expression (collectively also called proprioception).
- Gustatory system
- Olfactory system
Human sensory receptors are:
- Chemosensor
- Mechanoreceptor
- Nociceptor
- Photoreceptor
- Thermoreceptor
Diseases
Amblyopia
Anacusis
Astigmatism (eye)
Cataract
Color blindness
Deafness
Hyperopia
Disability-adjusted life year for sense organ diseases per 100,000 inhabitants in 2002.[3]
no data
less than 200
200-400
400-600
600-800
800-1000
1000-1200
1200-1400
1400-1600
1600-1800
1800-2000
2000-2300
more than 2300
See also
- Sensor
- Sensory neuroscience
- Wikibook "Sensory Systems"
References
- ^ John Krantz. Experiencing Sensation and Perception. "Chapter 1: What is Sensation and Perception?" pp. 1.6[1] Retrieved May 25, 2012.
- ^ Kolb & Whishaw: Fundamentals of Human Neuropsychology (2003)
- ^ "Mortality and Burden of Disease Estimates for WHO Member States in 2002" (xls). World Health Organization. 2002. http://www.who.int/entity/healthinfo/statistics/bodgbddeathdalyestimates.xls.
Human systems and organs
|
|
TA 2–4:
MS |
Skeletal system
|
- Bone
- Carpus
- Collar bone (clavicle)
- Thigh bone (femur)
- Fibula
- Humerus
- Mandible
- Metacarpus
- Metatarsus
- Ossicles
- Patella
- Phalanges
- Radius
- Skull (cranium)
- Tarsus
- Tibia
- Ulna
- Rib
- Vertebra
- Pelvis
- Sternum
- Cartilage
|
|
Joints
|
- Fibrous joint
- Cartilaginous joint
- Synovial joint
|
|
Muscular system
|
|
|
|
TA 5–11:
splanchnic/
viscus |
mostly
Thoracic
|
|
|
mostly
Abdominopelvic
|
Digestive system+
adnexa
|
- Mouth
- upper GI
- Oropharynx
- Laryngopharynx
- Esophagus
- Stomach
- lower GI
- Small intestine
- Appendix
- Colon
- Rectum
- Anus
- accessory
- Liver
- Biliary tract
- Pancreas
|
|
GU: Urinary system
|
- Kidney
- Ureter
- Bladder
- Urethra
|
|
GU: Reproductive system
|
- Female
- Uterus
- Vagina
- Vulva
- Ovary
- Placenta
- Male
- Scrotum
- Penis
- Prostate
- Testicle
- Seminal vesicle
|
|
|
Endocrine system
|
- Pituitary
- Pineal
- Thyroid
- Parathyroid
- Adrenal
- Islets of Langerhans
|
|
|
TA 12–16 |
Circulatory system
|
Cardiovascular system
|
- peripheral
- Artery
- Vein
- Lymphatic vessel
- Heart
|
|
Lymphatic system
|
|
|
|
Nervous system
|
|
|
Integumentary system
|
- Skin
- Subcutaneous tissue
- Breast
|
|
|
Blood
(Non-TA) |
|
|
General anatomy: systems and organs, regional anatomy, planes and lines, superficial axial anatomy, superficial anatomy of limbs
|
|
Nervous system: Sensory systems / senses (TA A15)
|
|
Special senses |
- Visual system/sight
- Auditory system/hearing
- Chemoreception
- Olfactory system/smell
- Gustatory system/taste
|
|
Touch |
- Pain
- Heat
- Balance
- Mechanoreception
- Proprioception
|
|
Other |
|
|
Sensory system – visual system – globe of eye (TA A15.2.1–6, TH 3.11.08.0-5, GA 10.1005)
|
|
Fibrous tunic (outer) |
Sclera |
- Episcleral layer
- Schlemm's canal
- Trabecular meshwork
|
|
Cornea |
- Limbus
- layers
- Epithelium
- Bowman's
- Stroma
- Descemet's
- Endothelium
|
|
|
|
Uvea/vascular tunic (middle) |
Choroid |
- Capillary lamina of choroid
- Bruch's membrane
- Sattler's layer
|
|
Ciliary body |
- Ciliary processes
- Ciliary muscle
|
|
Iris |
- Stroma
- Pupil
- Iris dilator muscle
- Iris sphincter muscle
|
|
|
Retina (inner) |
Layers |
- Inner limiting membrane
- Nerve fiber layer
- Ganglion cell layer
- Inner plexiform layer
- Inner nuclear layer
- Outer plexiform layer
- Outer nuclear layer
- External limiting membrane
- Layer of rods and cones
- Retinal pigment epithelium
|
|
Cells |
Photoreceptor cells (Cone cell, Rod cell) → (Horizontal cell) → Bipolar cell → (Amacrine cell) → Retina ganglion cell (Midget cell, Parasol cell, Bistratified cell, Giant retina ganglion cells, Photosensitive ganglion cell) → Diencephalon: P cell, M cell, K cell
Muller glia
|
|
Other |
|
|
|
Anterior segment |
- Anterior chamber
- Aqueous humour
- Posterior chamber
- Lens
- Capsule of lens
- Zonule of Zinn
|
|
Posterior segment |
|
|
Other |
- Ocular immune system
- Tapetum lucidum
- Keratocytes
|
|
|
anat (g/a/p)/phys/devp/prot
|
|
proc, drug (S1A/1E/1F/1L)
|
|
|
|
Sensory system: Auditory and Vestibular systems (TA A15.3, TH 3.11.09, GA 10.1029)
|
|
Outer ear |
- Pinna (Helix, Antihelix, Tragus, Antitragus, Incisura anterior auris, Earlobe)
- Ear canal
- Auricular muscles
- Eardrum (Umbo
- Pars flaccida)
|
|
Middle ear |
Tympanic cavity
|
- Labyrinthine wall/medial: Oval window
- Round window
- Secondary tympanic membrane
- Prominence of facial canal
- Promontory of tympanic cavity
- Mastoid wall/posterior: Mastoid cells
- Aditus to mastoid antrum
- Pyramidal eminence
- Tegmental wall/roof: Epitympanic recess
|
|
Ossicles
|
- Malleus (Neck of malleus, Superior ligament of malleus, Lateral ligament of malleus, Anterior ligament of malleus)
- Incus (Superior ligament of incus, Posterior ligament of incus)
- Stapes (Anular ligament of stapes)
|
|
Muscles
|
|
|
Eustachian tube
|
- Bony part of pharyngotympanic tube
- Cartilage of pharyngotympanic tube (Torus tubarius)
|
|
|
Inner ear/
(membranous labyrinth,
bony labyrinth) |
Auditory system
Cochlear labyrinth
|
General cochlea
|
- Scala vestibuli
- Helicotrema
- Scala tympani
- Modiolus
- Cochlear cupula
|
|
Perilymphatic space
|
- Perilymph
- Cochlear aqueduct
|
|
Cochlear duct /
scala media
|
- Reissner's/vestibular membrane
- Basilar membrane
- Endolymph
- Stria vascularis
- Spiral ligament
- Organ of Corti: Stereocilia
- Tectorial membrane
- Sulcus spiralis (externus, internus)
- Spiral limbus
|
|
Cells
|
- Claudius cell
- Boettcher cell
|
|
|
Vestibular system/
Vestibular labyrinth
|
- Static/translations/vestibule/endolymphatic duct: Utricle (Macula)
- Saccule (Macula, Endolymphatic sac)
- Kinocilium
- Otolith
- Vestibular aqueduct
- Canalis reuniens
- Kinetic/rotations: Semicircular canals (Superior, Posterior, Horizontal)
- Ampullary cupula
- Ampullae (Crista ampullaris)
|
|
|
|
|
Sensory system: Gustatory system (TA 15.4, GA 10.991)
|
|
Tongue |
Taste bud (Vallate papilla, Foliate papilla, Fungiform papilla) · Filiform papilla
|
|
Path |
medulla: Solitary tract (VII, IX, X) · Solitary nucleus (Gustatory nucleus)
pons: Central tegmental tract · Medial parabrachial nucleus (Hypothalamus, Amygdala)
thalamus: Ventral posteromedial nucleus
cerebrum: Posterior limb of internal capsule · Gustatory cortex
|
|
Other |
Basic tastes
|
|
|
|
Sensory system: Olfactory system / Olfaction / Rhinencephalon (TA 15.1, GA 10.992)
|
|
Olfactory epithelium |
Olfactory glands · Olfactory mucosa · Sustentacular cell
|
|
Olfactory nerve: 1° neuron |
Olfactory receptor neurons (Olfactory receptor) → Olfactory bulb (Glomeruli)
|
|
Olfactory nerve: 2° neuron |
Mitral cells → Olfactory tract → Olfactory trigone
|
|
Lateral olfactory stria/
Primary olfactory cortex |
Piriform cortex · EC-hippocampus system (Entorhinal cortex, Hippocampal formation) · Prepyriform area · Periamygdaloid cortex
Stria medullaris → Habenular nuclei
Amygdala → Stria terminalis → Hypothalamus
Medial forebrain bundle → Hypothalamus
|
|
Medial olfactory stria |
Anterior olfactory nucleus
|
|
|
|
Nervous system, receptors: somatosensory system (GA 10.1059)
|
|
Medial lemniscus |
- Touch/mechanoreceptors: Lamellar/Pacinian corpuscles – vibration
- Tactile/Meissner's corpuscles – light touch
- Merkel's discs – pressure
- Bulbous/Ruffini endings - stretch
- Free nerve endings – pain
- Hair cells
- Baroreceptor
- Proprioception: Golgi organ – tension/length
- Muscle spindle – velocity of change
- Intrafusal muscle fiber
- Nuclear chain fiber
- Nuclear bag fiber
|
|
Spinothalamic tract |
- Pain: Nociception and Nociceptors
- Temperature: Thermoreceptors
|
|
|
anat (h/r/t/c/b/l/s/a)/phys (r)/devp/prot/nttr/nttm/ntrp
|
noco/auto/cong/tumr, sysi/epon, injr
|
|
|
|
|
Systems and systems science
|
|
Systems categories |
- Systems theory
- Systems science
- Systems scientists
- Conceptual
- Physical
- Social
|
|
Systems |
- Biological
- Complex
- Complex adaptive
- Conceptual
- Coupled human-environment system
- Database management
- Dynamical
- Economical
- Ecosystem
- Formal
- Global Positioning System
- Human anatomy
- Information systems
- Legal systems
- Systems of measurement
- Metric system
- Multi-agent system
- Nervous system
- Nonlinearity
- Operating system
- Physical system
- Political system
- Sensory system
- Social system
- Solar System
- Systems art
|
|
Theoretical fields |
- Chaos theory
- Complex systems
- Control theory
- Cybernetics
- Earth system science
- Living systems
- Sociotechnical systems theory
- System dynamics
- Systems biology
- Systems ecology
- Systems engineering
- Systems neuroscience
- Systems psychology
- Systems science
- Systems theory
- World-systems theory
|
|
Systems scientists |
- Russell Ackoff
- William Ross Ashby
- Béla Bánáthy
- Gregory Bateson
- Stafford Beer
- Richard Bellman
- Ludwig von Bertalanffy
- Kenneth Boulding
- Murray Bowen
- West Churchman
- George Dantzig
- Heinz von Foerster
- Jay Wright Forrester
- Charles A S Hall
- George Klir
- Edward Lorenz
- Niklas Luhmann
- Humberto Maturana
- Margaret Mead
- Donella Meadows
- Mihajlo Mesarovic
- James Grier Miller
- Howard Odum
- Talcott Parsons
- Ilya Prigogine
- Anatol Rapoport
- Claude Shannon
- Qian Xuesen
- Francisco Varela
- Kevin Warwick
- Norbert Wiener
- Anthony Wilden
|
|