Stag beetles |
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Male Lamprima aurata |
Scientific classification |
Kingdom: |
Animalia |
Phylum: |
Arthropoda |
Class: |
Insecta |
Order: |
Coleoptera |
Suborder: |
Polyphaga |
Infraorder: |
Scarabaeiformia |
Superfamily: |
Scarabaeoidea |
Family: |
Lucanidae
Latreille, 1804 |
Subfamilies |
Aesalinae
Lampriminae
Lucaninae
Syndesinae
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Stag beetles are a group of about 1,200 species of beetle in the family Lucanidae, presently classified in four subfamilies[1] Some species grow up to over 12 cm (4.8 in), but most are about 5 cm (2 in).
Contents
- 1 Overview
- 2 Antler allometry
- 3 Taxonomy
- 4 References
- 5 External links
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Overview
The English name is derived from the large and distinctive mandibles found on the males of most species, which resemble the antlers of stags.
A well-known species in much of Europe is Lucanus cervus, referred to in some European countries (including United Kingdom) as "the" stag beetle (it is the largest terrestrial insect in Europe). Pliny the Elder noted that Nigidius called the stag beetle lucanus after the Italian region of Lucania where they were used as amulets. The scientific name of Lucanus cervus is this word, plus cervus, deer.
Male stag beetles use their jaws to wrestle each other for favoured mating sites in a manner that parallels the way stags fight over females. Fights may also be over food, such as tree sap and decaying fruits. Despite their often fearsome appearance they are not normally aggressive to humans.
Female stag beetles are usually smaller than the males, with smaller mandibles. As larvae, females can be distinguished from males by the presence of cream-coloured, fat ovaries visible through the skin around 2⁄3 down the larva's back.
The larvae feed for several years on rotting deciduous wood, growing through three larval stages until eventually pupating inside a pupal cell constructed from surrounding wood pieces and soil particles. In the final larval stage, "L3", the grubs of larger species, such as Prosopocoilus giraffa, may be the size of a human finger.
Antler allometry in
Metopodontus savagei
Antler allometry
Lucanidae express male-specific "weapon traits" (antler size) that often show size variations among individual males.This is termed scaling relationship or static allometry. Environmental conditions during development affect absolute weapon size but there are genetic factors.
Taxonomy
See Taxonomy of Lucanidae Currently the only means of identification of world species is the illustrated text by Mizunuma, T. and S. Nagai. 1994. The Lucanid Beetles of the World. Mushi-sha, Tokyo. 337 pp. based on the work Catalogue illustré des lucanides du globe in Encyclopédie Entomologique (series A 27: 1-223) by Robert Didier and Eugene Seguy.
References
- ^ Smith, A.B.T. (2006). A review of the family-group names for the superfamily Scarabaeoidea (Coleoptera) with corrections to nomenclature and a current classification. The Coleopterists Bulletin 60:144–204.
External links
- Data related to Lucanidae at Wikispecies
- Media related to Lucanidae at Wikimedia Commons
- Flickr Images
- Stag beetle info Research site containing lots of information on the stag beetle as well as information on current conservation schemes.
- M.J. Paulsen. "Annotated Checklist of the New World Lucanidae". http://www-museum.unl.edu/research/entomology/Guide/Scarabaeoidea/Lucanidae/Lucanidae-Catalog/LucanidaeC.htm. Checklist of New World stag beetles with links to pages with additional information and images.
- Taiwanese picture gallery
- Toma Libich. "Goliathus.com: gallery of Lucanidae". http://www.goliathus.com/en/en-lucanidae.html.
- "Gallery of Lucanidae". http://www.goliathus.cz/en/museum-lucanidae-2.html.
- J M Maes. "Biodiversity: section on Lucanidae with a world catalogue and a world bibliography". http://www.bio-nica.info.
- Asahinet Stag beetles on postage stamps and species illustrations.
- Lucanes du Monde Image rich French blog
- Images (portal)
- TOL
- The Lucanid (Stag) Beetles of the World Extra detailed specimen photobook 2009
- UNL Generic Guide to New World Scarabs- Lucanidae,
- UK Stag Beetle School Project A UK school project about Stag Beetles (collecting data on populations)
Extant Coleoptera families
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- Kingdom: Animalia
- Phylum: Arthropoda
- Class: Insecta
- Subclass: Pterygota
- Infraclass: Neoptera
- Superorder: Endopterygota
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Suborder Archostemata
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- Crowsoniellidae (Crowsoniella relicta)
- Cupedidae (reticulated beetles)
- Jurodidae (Sikhotealinia zhiltzovae)
- Micromalthidae (telephone-pole beetle)
- Ommatidae
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Suborder Adephaga
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- Amphizoidae (trout-stream beetles)
- Aspidytidae
- Carabidae (ground beetles)
- Dytiscidae (predaceous diving beetles)
- Gyrinidae (whirligig beetles)
- Haliplidae (crawling water beetles)
- Hygrobiidae
- Meruidae (Meru phyllisae)
- Noteridae (burrowing water beetles)
- Rhysodidae (wrinkled bark beetles)
- Trachypachidae (false ground beetles)
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Suborder Myxophaga
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- Hydroscaphidae (skiff beetles)
- Lepiceridae
- Sphaeriusidae
- Torridincolidae
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Suborder Polyphaga
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Bostrichiformia
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Bostrichoidea
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- Anobiidae (furniture beetles, death watch beetles, spider beetles)
- Bostrichidae (auger beetles)
- Dermestidae (skin beetles)
- Jacobsoniidae (Jacobson's beetles)
- Nosodendridae (wounded-tree beetles)
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Derodontoidea
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- Derodontidae (tooth-necked fungus beetles)
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Cucujiformia
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Chrysomeloidea
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- Cerambycidae (longhorn beetles)
- Chrysomelidae (leaf beetles)
- Disteniidae
- Megalopodidae
- Orsodacnidae
- Oxypeltidae
- Vesperidae
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Cleroidea
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- Acanthocnemidae (Acanthocnemus nigricans)
- Chaetosomatidae
- Cleridae (checkered beetles)
- Melyridae (soft-wing flower beetles)
- Phloiophilidae (Phloiophilus edwardsi)
- Phycosecidae
- Prionoceridae
- Trogossitidae (bark-gnawing beetles)
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Cucujoidea
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- Alexiidae
- Biphyllidae (false skin beetles)
- Boganiidae
- Bothrideridae (dry bark beetles)
- Byturidae (fruitworm beetles)
- Cavognathidae
- Cerylonidae (minute bark beetles)
- Coccinellidae (lady beetles, or God's cows)
- Corylophidae (minute fungus beetles)
- Cryptophagidae (silken fungus beetles)
- Cucujidae (flat bark beetles)
- Cyclaxyridae
- Discolomatidae
- Endomychidae (handsome fungus beetles)
- Erotylidae (pleasing fungus beetles)
- Helotidae
- Hobartiidae
- Kateretidae (short-winged flower beetles)
- Laemophloeidae (lined flat bark beetles)
- Lamingtoniidae (Lamingtonium binnaberrense)
- Latridiidae (minute brown scavenger beetles)
- Monotomidae (root-eating beetles)
- Myraboliidae
- Nitidulidae (sap beetles)
- Passandridae (parasitic flat bark beetles)
- Phalacridae (shining flower beetles)
- Phloeostichidae
- Propalticidae
- Protocucujidae
- Silvanidae (silvanid flat bark beetles)
- Smicripidae (palmetto beetles)
- Sphindidae (dry-fungus beetles)
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Curculionoidea
(weevils)
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- Anthribidae (fungus weevils)
- Attelabidae (leaf-rolling weevils)
- Belidae (primitive weevils)
- Brentidae (straight snout weevils, New York weevil)
- Caridae
- Curculionidae (true weevils, bark beetles, ambrosia beetles)
- Nemonychidae (pine flower weevils)
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Lymexyloidea
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- Lymexylidae (ship-timber beetles)
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Tenebrionoidea
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- Aderidae (ant-like leaf beetles)
- Anthicidae (ant-like flower beetles)
- Archeocrypticidae (cryptic fungus beetles)
- Boridae (conifer bark beetles)
- Chalcodryidae
- Ciidae (minute tree-fungus beetles)
- Melandryidae (false darkling beetles)
- Meloidae (blister beetles)
- Mordellidae (tumbling flower beetles)
- Mycetophagidae (hairy fungus beetles)
- Mycteridae (palm and flower beetles)
- Oedemeridae (false blister beetle)
- Perimylopidae, or Promecheilidae
- Prostomidae (jugular-horned beetles)
- Pterogeniidae
- Pyrochroidae (fire-coloured beetles)
- Pythidae (dead log bark beetles)
- Ripiphoridae (wedge-shaped beetles)
- Salpingidae (narrow-waisted bark beetles)
- Scraptiidae (false flower beetles)
- Stenotrachelidae (false longhorn beetles)
- Synchroidae (synchroa bark beetles)
- Tenebrionidae (darkling beetles)
- Tetratomidae (polypore fungus beetles)
- Trachelostenidae
- Trictenotomidae
- Ulodidae
- Zopheridae (ironclad beetles, cylindrical bark beetles)
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Elateriformia
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Buprestoidea
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- Buprestidae (jewel beetles, or metallic wood-boring beetles)
- Schizopodidae
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Byrrhoidea
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- Byrrhidae (pill beetles)
- Callirhipidae (cedar beetles)
- Chelonariidae (turtle beetles)
- Cneoglossidae
- Dryopidae (long-toed water beetles)
- Elmidae (riffle beetles)
- Eulichadidae (forest stream beetles)
- Heteroceridae (variegated mud-loving beetles)
- Limnichidae (minute mud beetles)
- Lutrochidae (travertine beetles)
- Psephenidae (water-penny beetles)
- Ptilodactylidae
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Dascilloidea
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- Dascillidae (soft bodied plant beetles)
- Rhipiceridae (cicada beetle, cicada parasite beetles)
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Elateroidea
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- Anischiidae
- Artematopodidae (soft-bodied plant beetles)
- Brachypsectridae (Texas beetles)
- Cantharidae (soldier beetles)
- Cerophytidae (rare click beetles)
- Drilidae
- Elateridae (click beetles)
- Eucnemidae (false click beetles)
- Lampyridae (fireflies)
- Lycidae (net-winged beetles)
- Omalisidae
- Omethidae (false fireflies)
- Phengodidae (glowworm beetles, long-lipped beetles)
- Plastoceridae (Plastocerus angulosus)
- Podabrocephalidae
- Rhagophthalmidae
- Rhinorhipidae (Rhinorhipus tamborinensis)
- Throscidae (false metallic wood-boring beetles)
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Scirtoidea
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- Clambidae
- Decliniidae (Declinia relicta)
- Eucinetidae (plate-thigh beetles)
- Scirtidae
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Scarabaeiformia
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Scarabaeoidea
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- Belohinidae (Belohina inexpectata)
- Bolboceratidae
- Ceratocanthidae
- Diphyllostomatidae (false stag beetles)
- Geotrupidae (dor beetles)
- Glaphyridae (bumble bee scarab beetles)
- Glaresidae (enigmatic scarab beetles)
- Hybosoridae (scavenger scarab beetles)
- Lucanidae (stag beetles)
- Ochodaeidae (sand-loving scarab beetles)
- Passalidae (betsy beetles)
- Pleocomidae (rain beetles)
- Scarabaeidae (scarabs)
- Trogidae (hide beetles)
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Staphyliniformia
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Histeroidea
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- Histeridae (clown beetles)
- Sphaeritidae (false clown beetles)
- Synteliidae
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Hydrophiloidea
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- Epimetopidae
- Georissidae (minute mud-loving beetles)
- Helophoridae
- Hydrochidae
- Hydrophilidae (water scavenger beetles)
- Spercheidae
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Staphylinoidea
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- Agyrtidae (primitive carrion beetles)
- Hydraenidae
- Leiodidae (round fungus beetles)
- Ptiliidae (feather-winged beetles)
- Scydmaenidae (ant-like stone beetles)
- Silphidae (carrion beetles)
- Staphylinidae (rove beetles)
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List of subgroups of the order Coleoptera
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