sense organ

感覚器

関

sensorium、sensory body

WordNet

1. the areas of the brain that process and register incoming sensory information and make possible the conscious awareness of the world
2. detect some circumstance or entity automatically; "This robot can sense the presence of people in the room"; "particle detectors sense ionization"
3. the faculty through which the external world is apprehended; "in the dark he had to depend on touch and on his senses of smell and hearing" (同)sensation, sentience, sentiency, sensory faculty
4. the meaning of a word or expression; the way in which a word or expression or situation can be interpreted; "the dictionary gave several senses for the word"; "in the best sense charity is really a duty"; "the signifier is linked to the signified" (同)signified
5. a general conscious awareness; "a sense of security"; "a sense of happiness"; "a sense of danger"; "a sense of self"
6. a natural appreciation or ability; "a keen musical sense"; "a good sense of timing"
7. comprehend; "I sensed the real meaning of his letter"
8. wind instrument whose sound is produced by means of pipes arranged in sets supplied with air from a bellows and controlled from a large complex musical keyboard (同)pipe_organ
9. a fully differentiated structural and functional unit in an animal that is specialized for some particular function
10. a government agency or instrument devoted to the performance of some specific function; "The Census Bureau is an organ of the Commerce Department"
11. a periodical that is published by a special interest group; "the organ of the communist party"
12. detected by instinct or inference rather than by recognized perceptual cues; "the felt presence of an intruder"; "a sensed presence in the room raised goosebumps on her arms"; "a perceived threat" (同)perceived
13. becoming aware of something via the senses (同)perception
14. a fractional monetary unit of Japan and Indonesia and Cambodia; equal to one hundredth of a yen or rupiah or riel

PrepTutorEJDIC

1. 〈C〉(五感による)『感覚』;(…の)感覚《+『of』+『名』》 / 〈U〉《しばしばa~》(…の)『感じ』,気持ち《+『of』+『名』》 / 〈U〉《a~,one's~》(…の)『理解』(『認識』)『力』,観念,センス《+『of』+『名』》 / 《one's senses》正常な感覚,正気 / 〈C〉《しばしば冠詩をつけないで》分別,良識 / 〈C〉(行動・発言・文などの)意義,意味(meaning)《+of(in)+名(do『ing』)》 / 〈U〉(…の)(全体的な)意向,動向《+『of』+『名』》 / …‘を'感ずる,‘に'気づく
2. 『オルガン』,パイプオルガン(pipe organ),リードオルガン(reed organ),電子オルガン(electronic organ),手回しオルガン(barrel organ) / (動植物の)『器官』 / (政党・会社などの)機関誌,機関紙 / 《しばしば複数形で》(行動・実施などの)組織,機関

Wikipedia preview

出典(authority):フリー百科事典『ウィキペディア（Wikipedia）』「2014/09/06 14:33:07」(JST)

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[Wiki en表示]

Sensory system
Typical sensory system: the visual system, illustrated by the classic Gray's FIG. 722– This scheme shows the flow of information from the eyes to the central connections of the optic nerves and optic tracts, to the visual cortex. Area V1 is the region of the brain which is engaged in vision.
Latin	organa sensuum
TA	A15.0.00.000
Anatomical terminology

The visual system and the somatosensory system are active even during resting state fMRI.

A sensory system is a part of the nervous system responsible for processing sensory information. A sensory system consists of sensory receptors, neural pathways, and parts of the brain involved in sensory perception. Commonly recognized sensory systems are those for vision, auditory (hearing), somatic sensation (touch), gustatory (taste), olfaction (smell) and vestibular (balance/movement). In short, senses are transducers from the physical world to the realm of the mind where we interpret the information, creating our perception of the world around us.

The receptive field is the specific part of the world to which a receptor organ and receptor cells respond. For instance, the part of the world an eye can see, is its receptive field; the light that each rod or cone can see, is its receptive field. Receptive fields have been identified for the visual system, auditory system and somatosensory system, so far.

Stimulus

Sensory systems code for four aspects of a stimulus; type (modality), intensity, location, and duration. Arrival time of a sound pulse and phase differences of continuous sound are used for localization of sound sources. Certain receptors are sensitive to certain types of stimuli (for example, different mechanoreceptors respond best to different kinds of touch stimuli, like sharp or blunt objects). Receptors send impulses in certain patterns to send information about the intensity of a stimulus (for example, how loud a sound is). The location of the receptor that is stimulated gives the brain information about the location of the stimulus (for example, stimulating a mechanoreceptor in a finger will send information to the brain about that finger). The duration of the stimulus (how long it lasts) is conveyed by firing patterns of receptors. These impulses are transmitted to the brain through afferent neurons.

Senses and receptors

While there is debate among neurologists as to the specific number of senses due to differing definitions of what constitutes a sense, Gautama Buddha and Aristotle classified five ‘traditional’ human senses which have become universally accepted: touch, taste, smell, sight, and hearing. Other senses that have been well-accepted in most mammals, including humans, include nociception, equilibrioception, kinaesthesia, and thermoception. Furthermore, some non-human animals have been shown to possess alternate senses, including magnetoception and electroreception.

Receptors

The initialization of sensation stems from the response of a specific receptor to a physical stimulus. The receptors which react to the stimulus and initiate the process of sensation are commonly characterized in four distinct categories: chemoreceptors, photoreceptors, mechanoreceptors, and thermoreceptors. All receptors receive distinct physical stimuli and transduce the signal into an electrical action potential. This action potential then travels along afferent neurons to specific brain regions where it is processed and interpreted.

Chemoreceptors

Main article: Chemoreceptor

Chemoreceptors, or chemosensors, detect certain chemical stimuli and transduce that signal into an electrical action potential. There are two primary types of chemoreceptors:

• Distance chemoreceptors are integral to receiving stimuli in the olfactory system through both olfactory receptor neurons and neurons in the vomeronasal organ.
• Direct chemoreceptors include the taste buds in the gustatory system as well as receptors in the aortic bodies which detect changes in oxygen concentration.

Photoreceptors

Main article: Photoreceptor cell

Photoreceptors are capable of phototransduction, a process which converts light (electromagnetic radiation) into, among other types of energy, a membrane potential. There are three primary types of photoreceptors: Cones are photoreceptors which respond significantly to color. In humans the three different types of cones correspond with a primary response to short wavelength (blue), medium wavelength (green), and long wavelength (yellow/red). Rods are photoreceptors which are very sensitive to the intensity of light, allowing for vision in dim lighting. The concentrations and ratio of rods to cones is strongly correlated with whether an animal is diurnal or nocturnal. In humans rods outnumber cones by approximately 20:1, while in nocturnal animals, such as the tawny owl, the ratio is closer to 1000:1. Ganglion Cells reside in the adrenal medulla and retina where they are involved in the sympathetic response. Of the ~1.3 million ganglion cells present in the retina, 1-2% are believed to be photosensitive.

Mechanoreceptors

Main article: Mechanoreceptor

Mechanoreceptors are sensory receptors which respond to mechanical forces, such as pressure or distortion. While mechanoreceptors are present in hair cells and play an integral role in the vestibular and auditory system, the majority of mechanoreceptors are cutaneous and are grouped into four categories:

• Slowly Adapting type 1 Receptors have small receptive fields and respond to static stimulation. These receptors are primarily used in the sensations of form and roughness.
• Slowly Adapting type 2 Receptors have large receptive fields and respond to stretch. Similarly to type 1, they produce sustained responses to a continued stimuli.
• Rapidly Adapting Receptors have small receptive fields and underlie the perception of slip.
• Pacinian Receptors have large receptive fields and are the predominant receptors for high frequency vibration.

Thermoreceptors

Main article: Thermoreceptor

Thermoreceptors are sensory receptors which respond to varying temperatures. While the mechanisms through which these receptors operate is unclear, recent discoveries have shown that mammals have at least two distinct types of thermoreceptors:

• The End-Bulb of Krause, or bulboid corpuscle, detects temperatures above body temperature
• Ruffini’s end organ detects temperatures below body temperature

Nociceptors

Main article: Nociceptor

Nociceptors respond to potentially damaging stimuli by sending signals to the spinal cord and brain. This process, called nociception, usually causes the perception of pain. They are found in internal organs as well as on the surface of the body. Nociceptors detect different kinds of damaging stimuli or actual damage. Those that only respond when tissues are damaged are known as "sleeping" or "silent" nociceptors.

• Thermal nociceptors are activated by noxious heat or cold at various temperatures.
• Mechanical nociceptors respond to excess pressure or mechanical deformation.
• Chemical nociceptors respond to a wide variety of chemicals, some of which are signs of tissue damage. They are involved in the detection of some spices in food.

Sensory cortex

All stimuli received by the receptors listed above are transduced to an action potential, which is carried along one or more afferent neurons towards a specific area of the brain. While the term sensory cortex is often used informally to refer to the somatosensory cortex, the term more accurately refers to the multiple areas of the brain at which senses are received to be processed. For the five traditional senses in humans, this includes the primary and secondary cortexes of the different senses: the somatosensory cortex, the visual cortex, the auditory cortex, the primary olfactory cortex, and the gustatory cortex.

Somatosensory cortex

Located in the parietal lobe, the somatosensory cortex is the primary receptive area for the sense of touch. This cortex is further divided into Brodmann areas 1, 2, and 3. Brodmann area 3 is considered the primary processing center of the somatosensory cortex as it receives significantly more input from the thalamus, has neurons highly responsive to somatosensory stimuli, and can evoke somatic sensations through electrical stimulation. Areas 1 and 2 receive most of their input from area 3.

Visual cortex

The visual cortex refers to the primary visual cortex, labeled V1 or Brodmann area 17, as well as the extrastriate visual cortical areas V2-V5. Located in the occipital lobe, V1 acts as the primary relay station for visual input, transmitting information to two primary pathways labeled the dorsal and ventral streams. The dorsal stream includes areas V2 and V5, and is used in interpreting visual ‘where’ and ‘how.’ The ventral stream includes areas V2 and V4, and is used in interpreting ‘what.’ Increases in Task-negative activity are observed in the ventral attention network, after abrupt changes in sensory stimuli, at the onset and offset of task blocks, and at the end of a completed trial.

Auditory cortex

Located in the temporal lobe, the auditory cortex is the primary receptive area for sound information. The auditory cortex is composed of Brodmann areas 41 and 42, also known as the anterior transverse temporal area 41 and the posterior transverse temporal area 42, respectively. Both areas act similarly and are integral in receiving and processing the signals transmitted from auditory receptors.

Primary olfactory cortex

Located in the temporal lobe, the primary olfactory cortex is the primary receptive area for olfaction, or smell. Unique to the olfactory and gustatory systems, at least in mammals, is the implementation of both peripheral and central mechanisms of action. The peripheral mechanisms involve olfactory receptor neurons which transduce a chemical signal along the olfactory nerve, which terminates in the olfactory bulb. The central mechanisms include the convergence of olfactory nerve axons into glomeruli in the olfactory bulb, where the signal is then transmitted to the anterior olfactory nucleus, the piriform cortex, the medial amygdala, and the entorhinal cortex, all of which make up the primary olfactory cortex.

Gustatory cortex

The gustatory cortex is the primary receptive area for taste, or gustation. The gustatory cortex consists of two primary structures: the anterior insula, located on the insular lobe, and the frontal operculum, located on the frontal lobe. Similarly to the olfactory cortex, the gustatory pathway operates through both peripheral and central mechanisms. Peripheral taste receptors, located on the tongue, soft palate, pharynx, and esophagus, transmit the received signal to primary sensory axons, where the signal is projected to the nucleus of the solitary tract in the medulla, or the gustatory nucleus of the solitary tract complex. The signal is then transmitted to the thalamus, which in turn projects the signal to several regions of the neocortex, including the gustatory cortex.

Modality

A stimulus modality (sensory modality) is a type of physical phenomenon that can be sensed. Examples are temperature, taste, sound, and pressure. The type of sensory receptor activated by a stimulus plays the primary role in coding the stimulus modality.

In the memory-prediction framework, Jeff Hawkins mentions a correspondence between the six layers of the cerebral cortex and the six layers of the optic tract of the visual system. The visual cortex has areas labelled V1, V2, V3, V4, V5, MT, IT, etc. Thus Area V1 mentioned below, is meant to signify only one class of cells in the brain, for which there can be many other cells which are also engaged in vision.

Hawkins lays out a scheme for the analogous modalities of the sensory system. Note that there can be many types of senses, some not mentioned here. In particular, for humans, there will be cells which can be labelled as belonging to V1, V2 A1, A2, etc.:

V1 (vision)

The human eye is the first element of a sensory system: in this case, vision, for the visual system.

Visual Area 1, or V1, is used for vision, via the visual system to the primary visual cortex.

ear

A1 (auditory - hearing)

Auditory Area 1, or A1, is for hearing, via the auditory system, the primary auditory cortex.

S1 (somatosensory - touch and proprioception)

Somatosensory Area 1, or S1, is for touch and proprioception in the somatosensory system. The somatosensory system feeds the Brodmann Areas 3, 1 and 2 of the primary somatosensory cortex. But there are also pathways for proprioception (via the cerebellum), and motor control (via Brodmann area 4). See also: S2 Secondary somatosensory cortex.

tongue

G1 (gustatory - taste)

Gustatory Area 1, or G1, is used for taste. Taste does not refer to the flavor of food, because the flavor of the food refers to the odor. The five qualities of taste include sourness, bitterness, sweetness, saltiness, and a recently discovered protein taste quality called umami.

O1 (olfactory - smell)

Olfactory Area 1, or O1, is used for smell. In contrast to vision and hearing, the olfactory bulbs are not cross-hemispheric; the right bulb connects to the right hemisphere and the left bulb connects to the left hemisphere.

Human sensory system

The Human sensory system consists of the following sub-systems:

• Visual system consists of the photoreceptor cells, optic nerve, and V1.
• Auditory system
• Somatosensory system consists of the receptors, transmitters (pathways) leading to S1, and S1 that experiences the sensations labelled as touch or pressure, temperature (warm or cold), pain (including itch and tickle), and the sensations of muscle movement and joint position including posture, movement, and facial expression (collectively also called proprioception).
• Gustatory system
• Olfactory system
• Vestibular system

Diseases

Disability-adjusted life year for sense organ diseases per 100,000 inhabitants in 2002.

  no data

  less than 200

  200-400

  400-600

  600-800

  800-1000

  1000-1200

  1200-1400

  1400-1600

  1600-1800

  1800-2000

  2000-2300

  more than 2300

• Amblyopia
• Anacusis
• Astigmatism (eye)
• Cataract
• Color blindness
• Deafness
• Hyperopia

See also

• Multisensory integration
• Neural adaptation
• Sensor
• Sensory neuroscience

References

External links

• Wikibook "Sensory Systems"

UpToDate Contents

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English Journal

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• Ribelles N1, Santonja A, Pajares B, Llácer C, Alba E.Author information 1Department of Medical Oncology, Hospital Universitario Virgen de la Victoria, Campus Teatinos s/n, 29010 Málaga, Spain. Electronic address: nuriaribelles@gmail.com.AbstractThe crucial event in the course of malignancies such as breast cancer is its metastatic spread from the primary tumor of origin to distant organs. The natural history of a tumor is determined by the expression of its genes, and in this sense, knowledge has advanced dramatically in recent decades. However, much less is known about the role that the patient plays in the behavior of a tumor. In this article, we review the evidence regarding the genetic background of the host in metastatic tumor dissemination, providing information from epidemiological studies as well as from animal models and human studies. Undoubtedly, the elucidation of possible interpersonal variability in susceptibility to developing metastases would significantly contribute to improve management of cancer patients.
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• Current opinion in psychiatry.Curr Opin Psychiatry.2014 Mar;27(2):128-37. doi: 10.1097/YCO.0000000000000041.
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• Muscle-bone interactions: basic and clinical aspects.

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Japanese Journal

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Related Links

• Sense organ - definition of sense organ by The Free Dictionary

sense organ n. A specialized organ or structure, such as the eye, ear, tongue, nose, or skin, where sensory neurons are concentrated and that functions as a receptor. Also called sensor. sense organ n 1. (Zoology) a structure in ...

• Sense Organ | Definition of sense organ by Merriam-Webster

Medical Definition of SENSE ORGAN: a bodily structure that receives a stimulus (as heat or sound waves) and is affected in such a manner as to initiate a wave of excitation in associated sensory nerve fibers which convey specific ...

Related Pictures

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★リンクテーブル★

リンク元	「sensorium」「sensory body」
関連記事	「sense」「sensing」「organ」

「sensorium」

　　[★]

感覚器

関

sense organ、sensory body

　

「sensory body」

　　[★]

感覚器

関

sense organ、sensorium

「sense」

　　[★]

• n.

• センス、意味、感覚

関

implication、imply、mean、meaning、purport、represent、semantic、sensation、sensory

　 　 　 　 　 　 　

「sensing」

　　[★]

• n.

• センシング

• adj.

• 感覚性の

関

sensory

　

「organ」

　　[★]

• n.

• 臓器、器官