canid

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Canidae

WordNet

1. dogs; wolves; jackals; foxes (同)family Canidae
2. stag beetles (同)family Lucanidae
3. pelicans (同)family Pelecanidae
4. the quantity contained in a can (同)canful
5. preserve in a can or tin; "tinned foods are not very tasty" (同)tin, put_up
6. airtight sealed metal container for food or drink or paint etc. (同)tin, tin_can
7. a buoy with a round bottom and conical top (同)can buoy

PrepTutorEJDIC

1. 《能力》『…することができる』　《可能性・推量》　《平叙文で》『…であり得る』　し得る,かもしれない　《否定文で》…のはずがない,ではあり得ない　《疑問文で》いったい…だろうか　《話》《許可・軽い命令》『…してもよい』　…してあげよう;《疑問文で》…してくれませんか
2. 『かん』(取っ手・ふた・口のついた金属・プラスチック製容器・コップなど) / 《米》(かん詰めの)『かん』,かん詰め(《英》 tin) / かん1杯(の…)《+『of』+『名』》 / …'を'かん詰めにする(また《英》 『tin』) / 《米俗》…'を'くびにする
3. canceled / cancellation / canon / canto

Wikipedia preview

出典(authority):フリー百科事典『ウィキペディア（Wikipedia）』「2015/07/03 11:40:34」(JST)

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Canids Temporal range: 39.75–0 Ma PreЄ Є O S D C P T J K Pg N Late Eocene – Recent
Major extant canid genera left-to-right, top-to-bottom: Canis, Cuon, Lycaon, Cerdocyon, Chrysocyon, Speothos, Vulpes, Nyctereutes, Otocyon and Urocyon
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Carnivora
Suborder:	Caniformia
Family:	Canidae G. Fischer de Waldheim, 1817
Genera and species
See text

The biological family Canidae /ˈkænɨdiː/ is a lineage of carnivorans that includes dogs, wolves, foxes, jackals, and many other extant and extinct dog-like mammals. A member of this family is called a canid (/ˈkænɨd/, /ˈkeɪnɨd/). The Canidae family is divided into two tribes: the Canini (dogs, wolves, jackals, and some south American "foxes") and the Vulpini (true foxes).

Canids have a long evolutionary history. In the Eocene, about 50 million years ago, the carnivorans split into two lineages, the caniforms (dog-like) and feliforms (cat-like). By the Oligocene, some ten million years later, the first proper canids had appeared and the family had split into three subfamilies, Hesperocyoninae, Borophaginae, and Caninae. Only the last of these has survived until the present day.

Canids are found on all continents except Antarctica and vary in size from the 2-m-long (6 ft 7 in) gray wolf to the 24-cm-long (9.4 in) fennec fox. The body forms of canids are similar, typically having long muzzles, upright ears, teeth adapted for cracking bones and slicing flesh, long legs, and bushy tails. They are mostly social animals, living together in family units or small groups and behaving cooperatively. Typically, only the dominant pair in a group breeds, and a litter of young is reared annually in an underground den. Canids communicate by scent signals and by vocalizations. One canid, the domestic dog (including the dingo), long ago entered into a partnership with humans and today remains one of the most widely kept domestic animals.

Phylogenetic relationships

Skulls of various Canid genera; Vulpes (corsac fox), Nyctereutes (raccoon dog), Cuon (dhole) and Canis (golden jackal)

Within the Canidae, the results of allozyme and chromosome analyses have previously suggested several phylogenetic divisions:

1. The wolf-like canids, (genus Canis, Cuon and Lycaon) including the dog, gray wolf (Canis lupus), red wolf (Canis rufus), eastern wolf (Canis lycaon), coyote (Canis latrans), golden jackal (Canis aureus), Ethiopian wolf (Canis simensis), black-backed jackal (Canis mesomelas), side-striped jackal (Canis adustus), dhole (Cuon alpinus), and African wild dog (Lycaon pictus).
2. The fox-like canids, which include the kit fox ("Vulpes aelox"), red fox (Vulpes vulpes), Cape fox (Vulpes chama), Arctic fox (Alopex lagopus), and fennec fox (Fennecus zerda).
3. The South American canids, including the bush dog (Speothos venaticus), hoary fox (Lycalopex uetulus), crab-eating fox (Cerdocyon thous) and maned wolf.
4. Various monotypic taxa, including the bat-eared fox (Otocyon megalotis), gray fox (Urocyon cinereoargenteus), and raccoon dog (Nycteruetes procyonoides).

Canidae family

Canidae Island fox Gray fox African wild dog Dhole Ethiopian wolf Coyote Gray wolf Dog Golden jackal Black-backed jackal Side-striped jackal Bush dog Maned wolf Hoary fox Darwin's fox Chilla Pampas fox Culpeo fox Sechuran fox Crab-eating fox Short-eared dog Bat-eared fox Raccoon dog Fennec fox Blanford's fox Cape fox Red fox Ruppell's fox Corsac fox Kit fox Arctic fox

Cladogram of Canidae from Lindblad-Toh (2005)

Recent DNA analysis shows that there are three monophyletic clades. Two of these, the South American canine group, which includes a number of "foxes", and the wolf group, together form the tribe Canini. The third clade is the "true fox" group, tribe Vulpini. Molecular data imply a North American origin of living Canidae some ten million years ago and an African origin of wolf-like canines (Canis, Cuon, and Lycaon), with the jackals being the most basal of this group. The South American clade is rooted by the maned wolf and bush dog, and the fox-like canids by the fennec fox and Blanford's fox. The grey fox and island fox are basal to the other clades, however this topological difference is not strongly supported.

Evolution

The Canidae includes a diverse group of some 34 species ranging in size from the maned wolf with its long limbs to the short-legged bush dog. Modern canids inhabit forests, tundra, savannahs and deserts throughout tropical and temperate parts of the world. The evolutionary relationships between the species have been studied in the past using morphological approaches but more recently, molecular studies have enabled the investigation of phylogenetic relationships. In some species, genetic divergence has been suppressed by the high level of gene flow between different populations and where the species have hybridized, large hybrid zones exist.

Eocene epoch

Carnivorans evolved from miacoids about 55 million years ago (Mya) during the late Paleocene. Some five million years later, the carnivorans split into two main divisions: caniforms (dog-like) and feliforms (cat-like). By 40 Mya, the first member of the dog family proper had arisen. Called Prohesperocyon wilsoni, its fossilized remains have been found in what is now the southwestern part of Texas. The chief features which identify it as a canid include the loss of the upper third molar (part of a trend toward a more shearing bite), and the structure of the middle ear which has an enlarged bulla (the hollow bony structure protecting the delicate parts of the ear). Prohesperocyon probably had slightly longer limbs than its predecessors, and also had parallel and closely touching toes which differ markedly from the splayed arrangements of the digits in bears.

The canid family soon subdivided into three subfamilies, each of which diverged during the Eocene: Hesperocyoninae (about 39.74-15 Mya), Borophaginae (about 34-2 Mya), and Caninae (about 34-0 Mya). Caninae is the only surviving subfamily and all present-day canids including wolves, foxes, coyotes, jackals, and domestic dogs belong to it. Members of each subfamily showed an increase in body mass with time, and some exhibited specialised hypercarnivorous diets that made them prone to extinction.

Evolution of the Canids

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An approximate timescale of key events in canid evolution.

For precise dates, see text.
Axis scale: millions of years ago.

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Oligocene epoch

By the Oligocene, all three subfamilies of canids (Hesperocyoninae, Borophaginae, and Caninae) had appeared in the fossil records of North America. The earliest and most primitive branch of the Canidae was the Hesperocyoninae lineage, which included the coyote-sized Mesocyon of the Oligocene (38-24 Mya). These early canids probably evolved for the fast pursuit of prey in a grassland habitat; they resembled modern civets in appearance. Hesperocyonines eventually became extinct in the middle Miocene. One of the early members of the Hesperocyonines, the genus Hesperocyon, gave rise to Archaeocyon and Leptocyon. These branches led to the borophagine and canine radiations.

Miocene epoch

Around 9–10 Mya during the Late Miocene, Canis, Urocyon, and Vulpes genera expanded from southwestern North America, where the canine radiation began. The success of these canines was related to the development of lower carnassials that were capable of both mastication and shearing. Around 8 Mya, the Beringian land bridge allowed members of the genus Eucyon a means to enter Asia and they continued on to colonise Europe.

Pliocene epoch

During the Pliocene, around 4–5 Mya, Canis lepophagus appeared in North America. This was small and sometimes coyote-like. Others were wolf-like in characteristics.Canis latrans (the coyote) is theorized to have descended from Canis lepophagus.

The formation of the Isthmus of Panama, about 3 Mya, joined South America to North America, allowing canids to invade South America, where they diversified. However the most recent common ancestor of the South American canids lived in North America some 4 Mya and the likelihood is that there were more than one incursion across the new land bridge. One of the resulting lineages consisted of the gray fox (Urocyon cinereoargentus) and the now extinct dire wolf (Canis dirus). The other lineage consisted of the so-called South American endemic species, the maned wolf (Chrysocyon brachyurus), the short-eared dog (Atelocynus microtis), the bush dog (Speothos venaticus), the crab-eating fox (Cerdocyon thous) and the South American foxes (Lycalopex spp.). The monophyly of this group has been established by molecular means.

Pleistocene epoch

During the Pleistocene, the North American wolf line appeared, with Canis edwardii, clearly identifiable as a wolf, and Canis rufus appeared, possibly a direct descendent of Canis edwardii. Around 0.8 Mya, Canis ambrusteri emerged in North America. A large wolf, it was found all over North and Central America, and was eventually supplanted by its descendant, the dire wolf, which then spread into South America during the late Pleistocene.

By 0.3 Mya, a number of subspecies of the gray wolf (Canis lupus) had developed and had spread throughout Europe and northern Asia. The gray wolf colonized North America during the late Rancholabrean era across the Bering land bridge, there being at least three separate invasions, with each one consisting of one or more different Eurasian gray wolf clades. MtDNA studies have shown that there are at least four extant C. lupus lineages. The dire wolf shared its habitat with the gray wolf but became extinct in a large-scale extinction event that occurred around 11,500 years ago. It may have been more of a scavenger than a hunter; its molars appear to be adapted for crushing bones and it may have died out as a result of the extinction of the large herbivorous animals on whose carcases it relied.

Characteristics

Comparative illustration of the paws of grey wolf, golden jackal and dhole by A. N. Komarov

Wild canids are found on every continent except Antarctica, and inhabit a wide range of different habitats, including deserts, mountains, forests, and grasslands. They vary in size from the fennec fox, which may be as little as 24 cm (9.4 in) in length and weigh 0.6 kg (1.3 lb), to the gray wolf, which may be up to 160 cm (5.2 ft) long, and can weigh up to 79 kg (174 lb). Only a few species are arboreal – the North American gray fox, the closely related Channel Island fox, and the raccoon dog habitually climb trees.

All canids have a similar basic form, as exemplified by the grey wolf, although the relative length of muzzle, limbs, ears and tail vary considerably between species. With the exceptions of the bush dog, raccoon dog, and some domestic breeds of Canis lupus, canids have relatively long legs and lithe bodies, adapted for chasing prey. The tails are bushy and the length and quality of the pelage varies with the season. The muzzle portion of the skull is much more elongated than that of the cat family. The zygomatic arches are wide, there is a transverse lambdoidal ridge at the rear of the cranium and in some species, a sagittal crest running from front to back. The bony orbits around the eye never form a complete ring and the auditory bullae are smooth and rounded.

All canids are digitigrade, meaning they walk on their toes. The tip of the nose is always naked, as are the cushioned pads on the soles of the feet. These latter consist of a single pad behind the tip of each toe and a more-or-less three-lobed central pad under the roots of the digits. Hairs grow between the pads and in the Arctic fox, the sole of the foot is densely covered with hair at some times of year. With the exception of the four-toed African hunting dog (Lycaon pictus), there are five toes on the forefeet but the pollex (thumb) is reduced and does not reach the ground. On the hind feet, there are four toes, but in some domestic dogs, a fifth vestigial toe, known as a dewclaw, is sometimes present but has no anatomical connection to the rest of the foot. The slightly curved nails are non-retractile and more or less blunt.

The penis in male canids is supported by a bone called the baculum. It also contains a structure at the base called the bulbus glandis which helps to create a copulatory tie during mating, locking the animals together for up to an hour. Young canids are born blind, with their eyes opening a few weeks after birth. All living canids (Caninae) have a ligament analogous to the nuchal ligament of ungulates used to maintain the posture of the head and neck with little active muscle exertion; this ligament allows them to conserve energy while running long distances following scent trails with their nose to the ground. However, based on skeletal details of the neck, at least some Borophaginae (such as Aelurodon) are believed to have lacked this ligament.

Dentition

Eurasian wolf skull

Most canids have 42 teeth, with a dental formula of: 3.1.4.23.1.4.3. The bush dog has only one upper molar with two below, the dhole has two above and two below, and the bat-eared fox has three or four upper molars and four lower ones. As in other members of Carnivora, the upper fourth premolar and lower first molar are adapted as carnassial teeth for slicing flesh, although the bat-eared fox differs in this respect, being largely insectivorous. The molar teeth are strong in most species, allowing the animals to crack open bone to reach the marrow. The deciduous, or baby teeth, formula in canids is 3.1.33.1.3, molars being completely absent.

Social behavior

Dholes attacking a sambar, Bandipur National Park

See also: Gray wolf § Behaviour, Dog behavior, Red fox § Behaviour and African Wild Dog § Social structure

Almost all canids are social animals and live together in groups. In general, they are territorial or have a home range and sleep in the open, using their dens only for breeding and sometimes in bad weather. In most foxes, and in many of the true dogs, a male and female pair work together to hunt and to raise their young. Gray wolves and some of the other larger canids live in larger groups called packs. African wild dogs have packs which may consist of twenty to forty animals, and packs of fewer than about seven individuals may be incapable of successful reproduction. Hunting in packs has the advantage that larger prey items can be tackled. Some species form packs or live in small family groups depending on the circumstances, including the type of available food. In most species, some individuals live on their own. Within a canid pack, there is a system of dominance so that the strongest, most experienced animals lead the pack. In most cases, the dominant male and female are the only pack members to breed.

Canids communicate with each other by scent signals, by visual clues and gestures, and by vocalizations such as growls, barks, and howls. In most cases, groups have a home territory from which they drive out other conspecifics. The territory is marked by leaving urine scent marks, which warn trespassing individuals. Social behaviour is also mediated by secretions from glands on the upper surface of the tail near its root and from the anal glands.

Reproduction

See also: Canine reproduction

Gray wolves and red foxes mating

A wild dog from Sri Lanka nursing her puppies

Canids as a group exhibit several reproductive traits that are uncommon among mammals as a whole. They are typically monogamous, provide paternal care to their offspring, have reproductive cycles with lengthy proestral and dioestral phases and have a copulatory tie during mating. They also retain adult offspring in the social group, suppressing the ability of these to breed while making use of the alloparental care they can provide to help raise the next generation of offspring.

During the proestral period, increased levels of oestradiol make the female attractive to the male. There is a rise in progesterone during the oestral phase and the female is now receptive. Following this, the level of oestradiol fluctuates and there is a lengthy dioestrous phase during which the female is pregnant. Pseudo-pregnancy frequently occurs in canids that have ovulated but failed to conceive. A period of anoestrus follows pregnancy or pseudo-pregnancy, there being only one oestral period during each breeding season. Small and medium-sized canids mostly have a gestation period of fifty to sixty days while larger species average sixty to sixty-five days. The time of year in which the breeding season occurs is related to the length of day, as has been demonstrated in the case of several species that have been translocated across the equator to the other hemisphere and experiences a six-month shift of phase. Domestic dogs and certain small canids in captivity may come into oestrus more frequently, perhaps because the photoperiod stimulus breaks down under conditions of artificial lighting.

The size of a litter varies,with from one to sixteen or more pups being born. The young are born small, blind and helpless and require a long period of parental care. They are kept in a den, most often dug into the ground, for warmth and protection. When the young begin eating solid food, both parents, and often other pack members, bring food back for them from the hunt. This is most often vomited up from the adult's stomach. Where such pack involvement in the feeding of the litter occurs, the breeding success rate is higher than is the case where females split from the group and rear their pups in isolation. Young canids may take a year to mature and learn the skills they need to survive. In some species, such as the African wild dog, male offspring usually remain in the natal pack, while females disperse as a group, and join another small group of the opposite sex to form a new pack.

Canids and humans

Traditional English fox hunt

Further information: domestic dog

See also: domesticated silver fox

One canid, the domestic dog, entered into a partnership with humans a long time ago. This partnership is documented as far back as 26,000 years ago, when the footprints of a young boy aged about eight to ten was found in Chauvet Cave in southern France, walking alongside what was identified as a large dog or wolf. The earliest recorded fossil of a dog was found to be around 36,000 years ago in Goyet Cave in Belgium. Even earlier, wolves were found fossilized in the same locations as humans at sites that date back 300,000 years, showing how far back humans and wolves had interactions with one another. The fact that wolves are pack animals with cooperative social structures may have been the reason that the relationship developed. Humans benefited from the canid's loyalty, cooperation, teamwork, alertness and tracking abilities while the wolf may have benefited from the use of weapons to tackle larger prey and the sharing of food. Humans and dogs may have evolved together. The bond between humans and dogs can be seen in the burial of dogs with their owners as early as 11,000 years ago in the Americas and 8,500 years ago in Europe.

Among canids, only the gray wolf has widely been known to prey on humans. Nonetheless, at least two records have coyotes killing humans, and two have golden jackals killing children. Human beings have trapped and hunted some canid species for their fur and, especially the gray wolf, coyote and the red fox, for sport. Canids such as the dhole are now endangered in the wild because of persecution, habitat loss, a depletion of ungulate prey species and transmission of diseases from domestic dogs.

Extant and recently extinct species

The genus Canis: Gray wolf, coyote, golden jackal, Ethiopian wolf, side-striped jackal and black-backed jackal

The genus Lycalopex: Culpeo, pampas fox, chilla and Darwin's fox

The genus Vulpes: red fox, Rüppell's fox, corsac fox, Bengal fox, Arctic fox, Blanford's fox, Cape fox and fennec fox

Dhole

African hunting dog

Short-eared dog

Maned wolf

Bush dog

Gray fox

Bat-eared fox

Raccoon dog

All extant species of family Canidae are in subfamily Caninae.

Subfamily Caninae

• True dogs – Tribe Canini
  • Genus Canis
    • Gray wolf, Canis lupus 2.723 Mya to present)
      • Domestic dog, Canis lupus familiaris
      • Dingo, most often classified as Canis lupus dingo (sometimes considered a separate taxon)
      • many other subspecies
    • Coyote, Canis latrans (also called prairie wolf)
    • Golden jackal, Canis aureus
    • Ethiopian wolf, Canis simensis (also called Abyssinian wolf, simien fox and simien jackal)
    • Side-striped jackal, Canis adustus
    • Black-backed jackal, Canis mesomelas
  • Genus Cuon
    • Dhole, Cuon alpinus or Canis alpinus (also called Asian wild dog)
  • Genus Lycaon
    • African wild dog, Lycaon pictus (also called African hunting dog)
  • Genus Atelocynus
    • Short-eared dog, Atelocynus microtis
  • Genus Cerdocyon
    • Crab-eating fox, Cerdocyon thous
  • Genus Dusicyon †
    • Falklands wolf, Dusicyon australis †
  • Genus Lycalopex (Pseudalopex)
    • Culpeo, Lycalopex culpaeus
    • Darwin's fox, Lycalopex fulvipes
    • South American gray fox, Lycalopex griseus
    • Pampas fox, Lycalopex gymnocercus
    • Sechura fox, Lycalopex sechurae
    • Hoary fox, Lycalopex vetulus
  • Genus Chrysocyon
    • Maned wolf, Chrysocyon brachyurus
  • Genus Speothos
    • Bush dog, Speothos venaticus
• True foxes – Tribe Vulpini
  • Genus Vulpes
    • Arctic fox, Vulpes lagopus
    • Red fox, Vulpes vulpes (1 Mya to present), including a domesticated silver fox
    • Swift fox, Vulpes velox
    • Kit fox, Vulpes macrotis
    • Corsac fox, Vulpes corsac
    • Cape fox, Vulpes chama
    • Pale fox, Vulpes pallida
    • Bengal fox, Vulpes bengalensis
    • Tibetan sand fox, Vulpes ferrilata
    • Blanford's fox, Vulpes cana
    • Rüppell's fox, Vulpes rueppelli
    • Fennec fox, Vulpes zerda
  • Genus Urocyon (2 Mya to present)
    • Gray fox, Urocyon cinereoargenteus
    • Island fox, Urocyon littoralis
    • Cozumel fox, Urocyon sp.
• Basal Caninae
  • Genus Otocyon (probably a vulpine close to Urocyon)
    • Bat-eared fox, Otocyon megalotis
  • Genus Nyctereutes
    • Raccoon dog, Nyctereutes procyonoides

Fluctuation of species within Canidae over 40 million years

Prehistoric Canidae

Except where otherwise stated, the following classification is based on a 1994 paper by Xiaoming Wang, curator of terrestrial mammals at the Natural History Museum of Los Angeles County on the systematics of the subfamily Hesperocyoninae, a 1999 paper by Wang, together with the zoologists Richard H. Tedford and Beryl E. Taylor on the subfamily Borophaginae, and a 2009 paper by Tedford, Wang and Taylor on the North American fossil Caninae.

Subfamily Caninae

• Tribe Canini
  • Genus Canis
    • Canis adoxus †
    • Canis ameghinoi †
    • Canis apolloniensis (1.1 mya) †
    • Canis armbrusteri (1.5 mya) †
    • Canis arnensis (3.4 Mya, †)
    • Canis cautleyi †
    • Canis cedazoensis (4.6 mya) †
    • Canis dirus (Dire wolf), (1.69 mya) †
    • Canis donnezani (4.0–3.1 Ma †, probably ancestor of wolves)
    • Canis edwardii (4.6 mya) †, first species of wolf in North America)
    • Canis (Eucyon) cipio (8.2 Mya †, probably first species of Canis genus)
    • Canis etruscus (3.4 Mya †)
    • Canis ferox (5 mya) †
    • Canis gezi †
    • Canis lepophagus (8 mya)†
    • Canis michauxi †
    • Canis mosbachensis (0.787 Mya †)
    • Canis nehringi †
  • Genus Cynotherium †
    • Sardinian dhole, Cynotherium sardous †
  • Genus Theriodictis (1.19 mya)†
    • Theriodictis platensis (1.8 Mya †)
    • Theriodictis tarijensis (1.8 Mya †)
    • Theriodictis (Canis) proplatensis (2.1 Ma † )
  • Genus Protocyon †
    • Protocyon orcesi †
    • Protocyon scagliarum †
    • Protocyon troglodytes †
  • Genus Dusicyon †
    • Dusicyon avus †
  • Genus Cerdocyon
    • Cerdocyon avius †
    • Cerdocyon ensenadensis †
  • Genus Speothos
    • Speothos pacivorus †
  • Genus Nurocyon †
    • Nurocyon chonokhariensis †
  • Genus Xenocyon †
    • Xenocyon falconeri (2.6 Mya †)
    • Xenocyon lycaonoides (1.69 mya) †
• Tribe Vulpini
  • Genus Vulpes (7 Mya to present)
    • Vulpes alopecoides (2.6 Mya †)
    • Vulpes cf. alopecoides (2.6 Mya †)
    • Vulpes cf. vulpes (0.1275 Mya †)
    • Vulpes galaticus (4.2 Mya †)
    • Vulpes riffautae (7 Mya †)
• Basal Caninae
  • Genus Nyctereutes (7.1 Mya to present)
    • Nyctereutes cf. donnezani (7.1 Mya †)
    • Nyctereutes cf. megamastoides (3.158 Mya †)
    • Nyctereutes donnezani (3.4 Mya †)
    • Nyctereutes megamostoides (2.6 Mya †)
    • Nyctereutes sinensis (3.4 Mya †)
• First Caninae
  • Genus Eucyon (8 Mya †)
    • Eucyon davisi (8.3 Mya †, probably ancestor of Canis)
    • Eucyon minor (8 Mya †)
    • Eucyon zhoui (8 Mya †)
    • Eucyon monticinensis(8 Mya †)
    • Eucyon odessanus †
  • Genus Leptocyon (24–16 Mya †)
    • Leptocyon vafer (16 Mya †)
    • Leptocyon vulpinus (24 Mya †)

Subfamily Borophaginae

† (Mya = million years ago) (million years = in existence)

• • Genus Archaeocyon (7 million years) †
    • Archaeocyon falkenbachi (10.2 million years) †
    • Archaeocyon leptodus (7 million years) †
    • Archaeocyon pavidus (7 million years) †
  • Genus Otarocyon (7.6 million years) †
    • Otarocyon cooki (4.5 million years) †
    • Otarocyon macdonaldi (0.6 million years) †
  • Genus Oxetocyon (2.5 million years) †
    • Oxetocyon cuspidatus (2.5 million years) †
  • Genus Rhizocyon (30 Mya)
    • Rhizocyon oregonensis (30 Mya)
• Tribe Phlaocyonini (27.2 million years) †
  • Genus Cynarctoides (16.7 million years) †
    • Cynarctoides acridens (11 million years) †
    • Cynarctoides emryi (4 million years) †
    • Cynarctoides gawnae (4 million years) †
    • Cynarctoides harlowi (4 million years) †
    • Cynarctoides lemur (30 Mya) †
    • Cynarctoides luskensis (4.2 million years) †
    • Cynarctoides roii (4.5 million years) †
  • Genus Phlaocyon (30–19 Mya)
    • Phlaocyon achoros
    • Phlaocyon annectens (22 Mya)
    • Phlaocyon latidens (30 Mya)
    • Phlaocyon leucosteus (22 Mya)
    • Phlaocyon mariae
    • Phlaocyon marslandensis (19 Mya)
    • Phlaocyon minor (30 Mya)
    • Phlaocyon multicuspus
    • Phlaocyon taylori
    • Phlaocyon yakolai (19 Mya)
• Tribe Borophagini (16.7 million years) †
  • Genus Cormocyon (10.2 million years) †
    • Cormocyon copei (10.2 million years) †
    • Cormocyon haydeni (4.2 million years) †
  • Genus Desmocyon (9 million years) †
    • Desmocyon matthewi (4.3 million years) †
    • Desmocyon thompsoni †
  • Genus Metatomarctus (4.3 million years) †
    • Metatomarctus canavus (4.3 million years) †
    • Metatomarctus sp. A (16 Mya)
    • Metatomarctus sp. B (16 Mya)
  • Genus Eulopocyon (18–16 Mya)
    • Eulopocyon brachygnathus (16 Mya)
    • Eulopocyon spissidens (18 Mya)
  • Genus Psalidocyon (16 Mya)
    • Psalidocyon marianae (16 Mya)
  • Genus Microtomarctus (4 million years) †
    • Microtomarctus conferta (17.67 million years) †
  • Genus Protomarctus (18 Mya)
    • Protomarctus optatus (18 Mya)
  • Genus Tephrocyon (16 Mya)
    • Tephrocyon rurestris (16 Mya)
  • Subtribe Cynarctina †
    • Genus Paracynarctus (6.7 million years) †
      • Paracynarctus kelloggi (6.7 million years) †
      • Paracynarctus sinclairi (7 million years) †
    • Genus Cynarctus (5.6 million years) †
      • Cynarctus crucidens (1.3 million years) †
      • Cynarctus galushai (2.7 million years) †
      • Cynarctus marylandica (2.3 million years) †
      • Cynarctus saxatilis (2.7 million years) †
      • Cynarctus voorhiesi (3.3 million years) †
  • Subtribe Aelurodontina (15 million years) †
    • Genus Tomarctus (7 million years) †
      • Tomarctus brevirostris (6.8 million years) †
      • Tomarctus hippophaga (7 million years) †
    • Genus Aelurodon (15.7 million years) †
      • Aelurodon asthenostylus (7 million years) †
      • Aelurodon ferox (6 mya) †
      • Aelurodon mcgrewi (2.7 mya) †
      • Aelurodon montanensis (2.7 mya) †
      • Aelurodon stirtoni (6 million years) †
      • Aelurodon taxoides (8.3 million years) †
  • Subtribe Borophagina (17 million years) †
    • Genus Paratomarctus (6 million years) †
      • Paratomarctus euthos (13 Mya)
      • Paratomarctus temerarius (16 Mya)
    • Genus Carpocyon (19.7 million years) †
      • Carpocyon compressus (2.7 million years) †
      • Carpocyon limosus (5 million years) †
      • Carpocyon robustus (3.3 million years) †
      • Carpocyon webbi (6 million years) †
    • Genus Protepicyon (16 Mya)
      • Protepicyon raki (16 Mya)
    • Genus Epicyon (2 million years) †
      • Epicyon aelurodontoides (5.4 million years) †
      • Epicyon haydeni (5.4 million years) †
      • Epicyon saevus (11.4 million years) †
    • Genus Borophagus (7 million years) †
      • Borophagus diversidens (5 Mya) †
      • Borophagus dudleyi (1.7 million years) †
      • Borophagus hilli (6.7 million years) †
      • Borophagus littoralis (0.6 million years) †
      • Borophagus orc (0.4 million years) †
      • Borophagus parvus (5.4 million years) †
      • Borophagus pugnator (8.3 million years) †
      • Borophagus secundus (8.3 million years) †

Subfamily Hesperocyoninae

† (Mya = million years ago)

• • Genus Cynodesmus (32–29 Mya)
    • Cynodesmus martini (29 Mya)
    • Cynodesmus thooides (32 Mya)
  • ?Genus Caedocyon
    • Caedocyon tedfordi
  • Genus Ectopocynus (32–19 Mya)
    • Ectopocynus antiquus (32 Ma)
    • Ectopocynus intermedius (29 Mya)
    • Ectopocynus siplicidens (19 Mya)
  • Genus Enhydrocyon (29–25 Mya)
    • Enhydrocyon basilatus (25 Mya)
    • Enhydrocyon crassidens (25 Mya)
    • Enhydrocyon pahinsintewkpa (29 Mya)
    • Enhydrocyon stenocephalus (29 Mya)
  • Genus Hesperocyon (39.74–34 Mya)
    • Hesperocyon coloradensis
    • Hesperocyon gregarius (37 Mya)
  • Genus Mesocyon (34–29 Mya)
    • Mesocyon brachyops (29 Mya)
    • Mesocyon coryphaeus (29 Mya)
    • Mesocyn temnodon
  • Genus Osbornodon (32–18 Mya)
    • Osbornodon brachypus
    • Osbornodon fricki (18 Mya)
    • Osbornodon iamonensis (21 Mya)
    • Osbornodon renjiei (33 Mya)
    • Osbornodon scitulus
    • Osbornodon sesnoni (32 Mya)
    • Osbornodon wangi
  • Genus Paraenhydrocyon (30–25 Mya)
    • Paraenhydrocyon josephi (30 Mya)
    • Paraenhydrocyon robustus (25 Mya)
  • Genus Philotrox (29 Mya)
    • Philotrox condoni (29 Mya)
  • Genus Prohesperocyon (36 Mya)
    • Prohesperocyon wilsoni (36 Mya)
  • Genus Sunkahetanka (29 Mya)
    • Sunkahetanka geringensis (29 Mya)

References

General references

• Xiaoming Wang, Richard H. Tedford, Mauricio Antón, Dogs: Their Fossil Relatives and Evolutionary History, New York : Columbia University Press, 2008; ISBN 978-0-231-13528-3

External links

• Canidae at the National Center for Biotechnology Information
• African Wild Dog Conservancy

UpToDate Contents

全文を閲覧するには購読必要です。 To read the full text you will need to subscribe.

• 1. エキノコックス症の治療および予防 treatment and prevention of echinococcosis

English Journal

• The complete mitochondrial genome of Chinese Shinjang wolf.

• Zhang H1, Zhang J, Chen L, Liu G.Author information 1College of Life Science, Qufu Normal University , Qufu , China.AbstractAbstract In this study, the complete mitochondrial genome of Chinese Shinjang wolf (Canis lupus desertorum) was sequenced for the first time using blood samples obtained from a female wolf captured from Altay in Shinjang, China. Sequence analysis showed that the genome structure was in accordance with other Canidae species and it contains 12S rRNA gene, 16S rRNA gene, 22 tRNA genes, 13 protein-coding genes and 1 control region.
• Mitochondrial DNA.Mitochondrial DNA.2014 Apr;25(2):106-8. doi: 10.3109/19401736.2013.786707. Epub 2013 May 8.
• Abstract In this study, the complete mitochondrial genome of Chinese Shinjang wolf (Canis lupus desertorum) was sequenced for the first time using blood samples obtained from a female wolf captured from Altay in Shinjang, China. Sequence analysis showed that the genome structure was in accordance wi
• PMID 23656292

• Neotropical echinococcosis: Second report of Echinococcus vogeli natural infection in its main definitive host, the bush dog (Speothos venaticus).

• do Carmo Pereira Soares M1, Souza de Souza AJ2, Pinheiro Malheiros A3, Marceliano Nunes H3, Almeida Carneiro L2, Moreira Alves M3, Farias da Conceição B3, Soares Gomes-Gouvêa M4, Marins Póvoa M5.Author information 1Seção de Hepatologia, Instituto Evandro Chagas/SVS/MS, CEP 66093-020, Belém, PA, Brazil. Electronic address: manoelsoares@iec.pa.gov.br.2Seção de Hepatologia, Instituto Evandro Chagas/SVS/MS, CEP 66093-020, Belém, PA, Brazil; Departamento de Patologia, Faculdade de Medicina Veterinária e Zootecnia, Universidade de São Paulo, CEP 05508-270, São Paulo, SP, Brazil.3Seção de Hepatologia, Instituto Evandro Chagas/SVS/MS, CEP 66093-020, Belém, PA, Brazil.4Laboratório de Gastroenterologia e Hepatologia Tropical, Instituto de Medicina Tropical, Faculdade de Medicina, Universidade de São Paulo, CEP 05403-000, São Paulo, SP, Brazil.5Seção de Parasitologia, Instituto Evandro Chagas/SVS/MS, CEP 67030-000, Ananindeua, PA, Brazil.AbstractThe bush dog (Speothos venaticus) acts as the natural definitive host in the life cycle of Echinococcus vogeli, the causative agent of polycystic hydatid disease, a zoonotic neglected disease in the South America. We report a case of natural infection by Echinococcus vogeli in a bush dog from the Brazilian Amazon, confirmed by the morphological and morphometric examination of adult parasites and their hooks obtained from the small intestine of the canid. Additionally, mitochondrial DNA sequence analysis corroborated these findings. This is the second report of natural infection by E. vogeli in a bush dog.
• Parasitology international.Parasitol Int.2014 Apr;63(2):485-7. doi: 10.1016/j.parint.2013.10.004. Epub 2013 Oct 19.
• The bush dog (Speothos venaticus) acts as the natural definitive host in the life cycle of Echinococcus vogeli, the causative agent of polycystic hydatid disease, a zoonotic neglected disease in the South America. We report a case of natural infection by Echinococcus vogeli in a bush dog from the Br
• PMID 24148288

• Isolation of viable Neospora caninum from brains of wild gray wolves (Canis lupus).

• Dubey JP1, Jenkins MC2, Ferreira LR2, Choudhary S2, Verma SK2, Kwok OC2, Fetterer R2, Butler E3, Carstensen M3.Author information 1United States Department of Agriculture, Agricultural Research Service, Beltsville Agricultural Research Center, Animal Parasitic Diseases Laboratory, Beltsville, MD 20705-2350, USA. Electronic address: Jitender.dubey@ars.usda.gov.2United States Department of Agriculture, Agricultural Research Service, Beltsville Agricultural Research Center, Animal Parasitic Diseases Laboratory, Beltsville, MD 20705-2350, USA.3Division of Wildlife, Minnesota Department of Natural Resources, 5463-C West Broadway, Forest Lake, MN 55025, USA.AbstractNeospora caninum is a common cause of abortion in cattle worldwide. Canids, including the dog and the dingo (Canis familiaris), the coyote (Canis latrans), and the gray wolf (Canis lupus) are its definitive hosts that can excrete environmentally resistant oocysts in the environment, but also can act as intermediate hosts, harboring tissue stages of the parasite. In an attempt to isolate viable N. caninum from tissues of naturally infected wolves, brain and heart tissue from 109 wolves from Minnesota were bioassayed in mice. Viable N. caninum (NcWolfMn1, NcWolfMn2) was isolated from the brains of two wolves by bioassays in interferon gamma gene knockout mice. DNA obtained from culture-derived N. caninum tachyzoites of the two isolates were analyzed by N. caninum-specific Nc5 polymerase chain reaction and confirmed diagnosis. This is the first report of isolation of N. caninum from tissues of any wild canid host.
• Veterinary parasitology.Vet Parasitol.2014 Mar 17;201(1-2):150-3. doi: 10.1016/j.vetpar.2013.12.032. Epub 2014 Jan 9.
• Neospora caninum is a common cause of abortion in cattle worldwide. Canids, including the dog and the dingo (Canis familiaris), the coyote (Canis latrans), and the gray wolf (Canis lupus) are its definitive hosts that can excrete environmentally resistant oocysts in the environment, but also can act
• PMID 24522164

Japanese Journal

• Overwintering strategy of wild free-ranging and enclosure-housed Japanese raccoon dogs ( Nyctereutes procyonoides albus )

• 北尾 直也,Fukui D,Hashimoto M,Osborne PG
• International Journal of Biometeorology 53(2), 159-165, 2009-03
• … http://dx.doi.org/10.1007/s00484-008-0199-7The raccoon dog, Nyctereutes procyonoides, is a canid with a passive overwintering strategy in northern Europe. …
• NAID 120003203376

• Canid-specific primers for molecular sexing using tissue or noninvasive samples

• SEDDON J. M.
• Conservation Genetics 6, 147-149, 2005
• NAID 30012717431

• Seasonal Physiology of the Wild Raccoon Dog (Nyctereutes procyonoides)(Physiology)

• Asikainen Juha,Mustonen Anne-Mari,Hyvarinen Heikki [他],Nieminen Petteri
• Zoological science 21(4), 385-391, 2004-04-25
• … The raccoon dog (Nyctereutes procyonoides) is a canid omnivore with autumnal fattening and winter sleep. …
• NAID 110006282409

Related Links

• Canid | Facebook

Canid. 103 likes. Canid, n. (KAY-nid). est. Winter, 2008, outside Washington, DC, USA in Bloomington, IN, as of Autumn, 2009 in Philadelphia, PA, as of Summer, 2013 Drones & beats from vocals, pedals, tapes, machines ...

• Canid - Definition and More from the Free Merriam-Webster ...

ca·nid noun \ ˈ ka-nəd, ˈ kā-\ Definition of CANID: any of a family (Canidae) of carnivorous animals that includes the wolves, jackals, foxes, coyote, and the domestic dog Origin of CANID New Latin Canidae, from Canis, type genus ...

• canid - Definition of canid - Dictionary definitions you can ...

canid definition by American Heritage Dictionary noun Any of various widely distributed carnivorous mammals of the family Canidae, which includes the foxes, wolves, dogs, jackals, and coyotes. Origin: From New Latin Canidae, , ...

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リンク元	「Canidae」「イヌ科」
拡張検索	「lercanidipine」「lercanidipine hydrochloride」「canid herpesvirus 1」
関連記事	「can」

「Canidae」

　　[★]

イヌ科

• ヌ科

関

canid、Canis familiaris、dog

　

「イヌ科」

　　[★]

ラ

Canidae、canid

関

イヌ

「lercanidipine」

　　[★]

レルカニジピン

関

lercanidipine hydrochloride

「lercanidipine hydrochloride」

　　[★]

• 塩酸レルカニジピン

関

lercanidipine

「canid herpesvirus 1」

　　[★]

• イヌヘルペスウイルス1型

「can」

　　[★]

• aux.

• (過去could)
• (可能)～できる、～しようと思えばできる、～する能力がある。
• (推量)～する可能性がある、～することがあり得る、～する/になることがある、～しない/にならないとも限らない。
• (許可)～してもよい、～しても差し支えない。

関

able、feasible、possible