二倍体　　　

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of a cell or organism having two sets of chromosomes or twice the haploid number; "diploid somatic cells"
(genetics) an organism or cell having the normal amount of DNA per cell; i.e., two sets of chromosomes or twice the haploid number
the condition of being diploid

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(生物の染色体が)二倍性の,倍数の / 二倍体(半数染色体を二組持つ生物体・組織)

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出典(authority):フリー百科事典『ウィキペディア（Wikipedia）』「2017/04/05 21:37:15」(JST)

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Diploid cells have twice the amount of chromosomes as haploid cells.

Ploidy is the number of sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes.

Cells are described according to the number of sets present (the ploidy level): monoploid (1 set), diploid (2 sets), triploid (3 sets), tetraploid (4 sets), pentaploid (5 sets), hexaploid (6 sets), heptaploid^[1] or septaploid^[2] (7 sets), etc. The generic term polyploid is used to describe cells with three or more sets of chromosomes (triploid or a higher ploidy).

Humans are diploid organisms, carrying two complete sets of chromosomes: one set of 23 chromosomes from their father and one set of 23 chromosomes from their mother. The two sets combined provide a full complement of 46 chromosomes. This total number of chromosomes is called the chromosome number. When a species has a varying chromosome number, e.g. a diploid and tetraploid form, the chromosome number is called diploid number in the diploid form, and tetraploid number in the tetraploid form.

The number of chromosomes found in a single complete set of chromosomes is called the monoploid number (x). The haploid number (n) is unique to gametes (sperm or egg cells), and refers to the total number of chromosomes found in a gamete, which under normal conditions is half the total number of chromosomes in a somatic cell.

The haploid number for humans (half of 46) is 23; and the monoploid number equals 46 divided by the ploidy level of 2, which is also 23. When a human germ cell undergoes meiosis the two sets of 23 chromosomes are split in half to form gametes. After fusion of a male and a female gamete (fertilization) both containing 1 set of 23 chromosomes, the resulting zygote has 46 chromosomes: 2 sets of 23 chromosomes (22 autosomes, and 1 allosome).

The common potato (Solanum tuberosum) is an example of a tetraploid organism, carrying four sets of chromosomes. The potato plant inherits two sets of 12 chromosomes from the pollen parent, and two sets of 12 chromosomes from the ovule parent. The four sets combined provide a full complement of 48 chromosomes. The haploid number (half of 48) is 24. The monoploid number equals the chromosome number divided by the ploidy level: 48 chromosomes in total divided by a ploidy level of 4 equals a monoploid number of 12.

The commercial common potato crop is propagated vegetatively (asexual reproduction through mitosis),^[3] in which case new individuals are produced from a single parent, without the involvement of gametes and fertilization, and all the offspring are genetically equal to each other and to the parent.

Variation in euploidy for organisms with x=11
Ploidy	Nr of chromosomes	Somatic	Gametic
Diploid	2x=22	2n=22	n=11
Tetraploid	4x=44	2n=44	n=22
Hexaploid	6x=66	2n=66	n=33
Octoploid	8x=88	2n=88	n=44

Because the chromosome number is generally reduced only by the specialized process of meiosis, the somatic cells of the body inherit and maintain the chromosome number of the zygote. However, in many situations somatic cells double their copy number by means of endoreduplication as an aspect of cellular differentiation. For example, the hearts of two-year-old children contain 85% diploid and 15% tetraploid nuclei, but by 12 years of age the proportions become approximately equal, and adults examined contained 27% diploid, 71% tetraploid and 2% octaploid nuclei.^[4]

When a germ cell with an uneven number of chromosomes undergoes meiosis, the chromosomes can't be evenly divided between two cells resulting in aneuploid gametes. Triploid organisms for instance are usually sterile. Because of this, triploidy is a common way of making seedless fruit such as bananas and watermelons. If the fertilization of human gametes results in 3 sets of chromosomes the condition is called triploid syndrome.

1 Etymology
2 Case studies
3 Haploid and monoploid
4 Diploid
5 Homoploid
6 Zygoidy and azygoidy
7 Polyploidy
8 Variable or indefinite ploidy
9 Polyploidy in bacteria and archaea
10 Mixoploidy
11 Dihaploidy and polyhaploidy
12 Euploidy
13 Possible adaptive and ecological significance of variation in ploidy
14 References
15 Bibliography
16 External links

Etymology

The term ploidy is a back-formation from haploidy and diploidy. Ploid is a combination of Ancient Greek -παλτος (-paltos), -πλος (-plos), -πλόος (-plóos, "fold"), and -oid from Ancient Greek -ειδής (-eidḗs), -οειδής (-oeidḗs), from εἶδος (eîdos, “form, likeness”). The Greek word ἁπλόος haplóos means "no fold", and διπλόος diplóos means "diplo" or "two-fold". Diploid therefore means resembling or having the likeness of diplo (compare with humanoid "resembling a human").

Eduard Strasburger, coined the terms haploid and diploid in 1905:

Schließlich wäre es vielleicht erwünscht, wenn den Bezeichnungen Gametophyt und Sporophyt, die sich allein nur auf Pflanzen mit einfacher und mit doppelter Chromosomenzahl anwenden lassen, solche zur Seite gestellt würden, welche auch für das Tierreich passen. Ich erlaube mir zu diesem Zwecke die Worte Haploid und Diploid, bezw. haploidische und diploidische Generation vorzuschlagen.^[5]^[6]

David Haig suggests that Strasburger based the terms on Weismann's conception of the id (or germ plasm),^[7]^[8] hence haplo-id and diplo-id. The two terms were borrowed from German through William Henry Lang's 1908 translation of an 1906 textbook by Strasburger and colleagues.^[9]^{[citation needed]}

Technically, ploidy refers to the nucleus. Though at times authors may report the total ploidy of all nuclei present within the cell membrane of a syncytium,^[10] usually the ploidy of the nuclei present will be described. For example, a fungal dikaryon with two haploid nuclei is distinguished from the diploid in which the chromosomes share a nucleus and can be shuffled together.^[11] Nonetheless, because in most situations there is only one nucleus, it is commonplace to speak of the ploidy of a cell.

Case studies

It is also possible on rare occasions for the ploidy to increase in the germline, which can result in polyploid offspring and ultimately polyploid species. This is an important evolutionary mechanism in both plants and animals.^[12] As a result, it becomes desirable to distinguish between the ploidy of a species or variety as it presently breeds and that of an ancestor. The number of chromosomes in the ancestral (non-homologous) set is called the monoploid number (x), and is distinct from the haploid number (n) in the organism as it now reproduces. Both numbers n, and x, apply to every cell of a given organism.^{[citation needed]}

Common wheat is an organism where x and n differ. It has six sets of chromosomes, two sets from each of three different diploid species that are its distant ancestors. The somatic cells are hexaploid, with six sets of chromosomes, 2n = 6x = 42. The gametes are haploid for their own species, but triploid, with three sets of chromosomes, by comparison to a probable evolutionary ancestor, einkorn wheat. The monoploid number x = 7, and the haploid number n = 21.^{[citation needed]}

Tetraploidy (four sets of chromosomes, 2n = 4x) is common in plants, and also occurs in amphibians, reptiles, and insects. For example, species of Xenopus (African toads) form a ploidy series, featuring diploid (X.tropicalis, 2n=20), tetraploid (for example X.laevis, 4n=36), octaploid (for example X.wittei, 8n=72) and dodecaploid (for example X.ruwenzoriensis, 12n=108) species.^[13]

Over evolutionary time scales in which chromosomal polymorphisms accumulate, these changes become less apparent by karyotype - for example, humans are generally regarded as diploid, but the 2R hypothesis has confirmed two rounds of whole genome duplication in early vertebrate ancestors.

Ploidy can also differ with life cycle.^[14]^[15] In some insects it differs by caste. In humans, only the gametes are haploid, but in the Australian bulldog ant, Myrmecia pilosula, a haplodiploid species, haploid individuals of this species have a single chromosome, and diploid individuals have two chromosomes.^[16] In Entamoeba, the ploidy level varies from 4n to 40n in a single population.^[17] Alternation of generations occurs in many plants.

Some studies suggest that selection is more likely to favor diploidy in host species and haploidy in parasite species.^[18]

Haploid and monoploid

1.Haploid organism are on the left and Diploid organism on the right. 2.This is an haploid egg carrying the dominant purple gene. 3.This is a haploid sperm carrying the recessive blue gene. 4.This is a diploid sperm carrying the recessive blue gene. 5.This is an diploid egg carrying the dominant purple gene. 6.This is the short lived diploid state of haploid organisms. 7.This is the first stage of a zygote which has just been fertilized by a sperm. 8.The spores released by the diploid structure either express the mothers dominate gene or the fathers recessive gene. 9. The baby's cells express the dominant or recessive.

The nucleus of a eukaryotic cell is haploid if it has a single set of chromosomes, each one not being part of a pair. By extension a cell may be called haploid if its nucleus is haploid, and an organism may be called haploid if its body cells (somatic cells) are haploid. The number of chromosomes in a single set is called the haploid number, given the symbol n.

Gametes (sperm and ova) are haploid cells. The haploid gametes produced by most organisms combine to form a zygote with n pairs of chromosomes, i.e. 2n chromosomes in total. The chromosomes in each pair, one of which comes from the sperm and one from the egg, are said to be homologous. Cells and organisms with pairs of homologous chromosomes are called diploid. For example, most animals are diploid and produce haploid gametes. During meiosis, sex cell precursors have their number of chromosomes halved by randomly "choosing" one member of each pair of chromosomes, resulting in haploid gametes. Because homologous chromosomes usually differ genetically, gametes usually differ genetically from one another.^{[citation needed]}

All plants and many fungi and algae switch between a haploid and a diploid state, with one of the stages emphasized over the other. This is called alternation of generations. Most fungi and algae are haploid during the principal stage of their lifecycle, as are plants like mosses. Most animals are diploid, but male bees, wasps, and ants are haploid organisms because they develop from unfertilized, haploid eggs.

In some cases there is evidence that the n chromosomes in a haploid set have resulted from duplications of an originally smaller set of chromosomes. This "base" number – the number of apparently originally unique chromosomes in a haploid set – is called the monoploid number,^[19] also known as basic or cardinal number,^[20] or fundamental number.^[21]^[22] As an example, the chromosomes of common wheat are believed to be derived from three different ancestral species, each of which had 7 chromosomes in its haploid gametes. The monoploid number is thus 7 and the haploid number is 3 × 7 = 21. In general n is a multiple of x. The somatic cells in a wheat plant have six sets of 7 chromosomes: three sets from the egg and three sets from the sperm which fused to form the plant, giving a total of 42 chromosomes. As a formula, for wheat 2n = 6x = 42, so that the haploid number n is 21 and the monoploid number x is 7. The gametes of common wheat are considered to be haploid, since they contain half the genetic information of somatic cells, but they are not monoploid, as they still contain three complete sets of chromosomes (n = 3x).^[23]

In the case of wheat, the origin of its haploid number of 21 chromosomes from three sets of 7 chromosomes can be demonstrated. In many other organisms, although the number of chromosomes may have originated in this way, this is no longer clear, and the monoploid number is regarded as the same as the haploid number. Thus in humans, x = n = 23.

Diploid

Diploid cells have two homologous copies of each chromosome, usually one from the mother and one from the father. Nearly all mammals are diploid organisms (the tetraploid (four sets) plains viscacha rats (Tympanoctomys barrerae) and golden vizcacha rat (Pipanacoctomys aureus)^[24] are the only known exceptions as of 2004^[25]), although all individuals have some small fraction of cells that display polyploidy. Human diploid cells have 46 chromosomes (the somatic number, 2n) and human haploid gametes (egg and sperm) have 23 chromosomes (n). Retroviruses that contain two copies of their RNA genome in each viral particle are also said to be diploid. Examples include human foamy virus, human T-lymphotropic virus, and HIV.^[26]

Homoploid

"Homoploid" means "at the same ploidy level", i.e. having the same number of homologous chromosomes. For example, homoploid hybridization is hybridization where the offspring have the same ploidy level as the two parental species. This contrasts with a common situation in plants where chromosome doubling accompanies, or happens soon after hybridization. Similarly, homoploid speciation contrasts with polyploid speciation.^{[citation needed]}

Zygoidy and azygoidy

Zygoidy is the state where the chromosomes are paired and can undergo meiosis. The zygoid state of a species may be diploid or polyploid.^[27]^[28] In the azygoid state the chromosomes are unpaired. It may be the natural state of some asexual species or may occur after meiosis. In diploid organisms the azygoid state is monoploid. (see below for dihaploidy)

Polyploidy

Polyploidy is the state where all cells have multiple sets of chromosomes beyond the basic set, usually 3 or more. Specific terms are triploid (3 sets), tetraploid (4 sets), pentaploid (5 sets), hexaploid (6 sets), heptaploid^[1] or septaploid^[2] (7 sets) octoploid (8 sets), nonaploid (9 sets), decaploid (10 sets), undecaploid (11 sets), dodecaploid (12 sets), tridecaploid (13 sets), tetradecaploid (14 sets) etc.^[29]^[30]^[31]^[32] Some higher ploidies include hexadecaploid (16 sets), dotriacontaploid (32 sets), and tetrahexacontaploid (64 sets),^[33] though Greek terminology may be set aside for readability in cases of higher ploidy (such as "16-ploid").^[31] Polytene chromosomes of plants and fruit flies can be 1024-ploid.^[34]^[35] Ploidy of systems such as the salivary gland, elaiosome, endosperm, and trophoblast can exceed this, up to 1048576-ploid in the silk glands of the commercial silkworm Bombyx mori.^[10]

The chromosome sets may be from the same species or from closely related species. In the latter case, these are known as allopolyploids (or amphidiploids, which are allopolyploids that behave as if they were normal diploids). Allopolyploids are formed from the hybridization of two separate species. In plants, this probably most often occurs from the pairing of meiotically unreduced gametes, and not by diploid–diploid hybridization followed by chromosome doubling.^[36] The so-called Brassica triangle is an example of allopolyploidy, where three different parent species have hybridized in all possible pair combinations to produce three new species.

Polyploidy occurs commonly in plants, but rarely in animals. Even in diploid organisms, many somatic cells are polyploid due to a process called endoreduplication where duplication of the genome occurs without mitosis (cell division).

The extreme in polyploidy occurs in the fern genus Ophioglossum, the adder's-tongues, in which polyploidy results in chromosome counts in the hundreds, or, in at least one case, well over one thousand.

It is also possible for polyploid organisms to revert to lower ploidy by means of haploidisation.

Variable or indefinite ploidy

Depending on growth conditions, prokaryotes such as bacteria may have a chromosome copy number of 1 to 4, and that number is commonly fractional, counting portions of the chromosome partly replicated at a given time. This is because under exponential growth conditions the cells are able to replicate their DNA faster than they can divide.

In ciliates, the macronucleus is called ampliploid, because only part of the genome is amplified.^[37]

Polyploidy in bacteria and archaea

Polyploidy is a characteristic of the bacterium Deinococcus radiodurans ^[38] and of the archaeon Halobacterium salinarum.^[39] These two species are highly resistant to ionizing radiation and desiccation, conditions that induce DNA double-strand breaks.^[40]^[41] This resistance appears to be due to efficient homologous recombinational repair.

Mixoploidy

Mixoploidy refers to the presence of two cell lines, one diploid and one polyploid, coexisting within the same organism. Though polyploidy in humans is not viable, mixoploidy has been found in live adults and children.^[42] There are two types: diploid-triploid mixoploidy, in which some cells have 46 chromosomes and some have 69,^[43] and diploid-tetraploid mixoploidy, in which some cells have 46 and some have 92 chromosomes. It is a major topic of cytology.

Dihaploidy and polyhaploidy

Dihaploid and polyhaploid cells are formed by haploidisation of polyploids, i.e., by halving the chromosome constitution.

Dihaploids (which are diploid) are important for selective breeding of tetraploid crop plants (notably potatoes), because selection is faster with diploids than with tetraploids. Tetraploids can be reconstituted from the diploids, for example by somatic fusion.

The term “dihaploid” was coined by Bender^[44] to combine in one word the number of genome copies (diploid) and their origin (haploid). The term is well established in this original sense,^[45]^[46] but it has also been used for doubled monoploids or doubled haploids, which are homozygous and used for genetic research.^[47]

Euploidy

Euploidy is the state of a cell or organism having one or more than one set of the same set of chromosomes, possibly excluding the sex-determining chromosomes. For example, most human cells have 2 of each of the 23 homologous monoploid chromosomes, for a total of 46 chromosomes. A human cell with an extra set out of the 23 normal ones would be considered euploid. Euploid karyotypes would consequentially be a multiple of the haploid number, which in humans is 23. Aneuploidy is the state where one or more chromosomes of a normal set are missing or present in more than their usual number of copies. Unlike euploidy, aneuploid karyotypes will not be a multiple of the haploid number. In humans, examples of anueploidy include having a single extra chromosome (such as Down syndrome), or missing a chromosome (such as Turner syndrome). Aneuploid karyotypes are given names with the suffix -somy (rather than -ploidy, used for euploid karyotypes), such as trisomy and monosomy.

Possible adaptive and ecological significance of variation in ploidy

A study comparing the karyotypes of endangered or invasive plants with those of their relatives found that being polyploid as opposed to diploid is associated with a 14% lower risk of being endangered, and a 20% greater chance of being invasive.^[48] Polyploidy may be associated with increased vigor and adaptability.^[49]

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Bibliography

Griffiths, A. J. et al. 2000. An introduction to genetic analysis, 7th ed. W. H. Freeman, New York ISBN 0-7167-3520-2

External links

Some eukaryotic genome-scale or genome size databases and other sources which may contain the ploidy of many organisms:

Animal genome size database
Plant genome size database
Fungal genome size database
Protist genome-scale database of Ensembl Genomes
Nuismer S.; Otto S.P. (2004). "Host-parasite interactions and the evolution of ploidy". Proc. Natl. Acad. Sci. USA. 101: 11036–11039. doi:10.1073/pnas.0403151101. (Supporting Data Set, with information on ploidy level and number of chromosomes of several protists)
Chromosome number and ploidy mutations YouTube tutorial video

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5. 巨核球の生物学と血小板の産生 megakaryocyte biology and the production of platelets

English Journal

Low recombination rates in sexual species and sex-asex transitions.

Haag CR1, Theodosiou L2, Zahab R3, Lenormand T3.
Philosophical transactions of the Royal Society of London. Series B, Biological sciences.Philos Trans R Soc Lond B Biol Sci.2017 Dec 19;372(1736). pii: 20160461. doi: 10.1098/rstb.2016.0461.
PMID 29109224

What drives the evolution of condition-dependent recombination in diploids? Some insights from simulation modelling.

Rybnikov SR1, Frenkel ZM2, Korol AB3.
Philosophical transactions of the Royal Society of London. Series B, Biological sciences.Philos Trans R Soc Lond B Biol Sci.2017 Dec 19;372(1736). pii: 20160460. doi: 10.1098/rstb.2016.0460.
PMID 29109223

Clinical Experience with the BCL2-inhibitor Venetoclax in Combination Therapy for Relapsed and Refractory Acute Myeloid Leukemia and Related Myeloid Malignancies.

DiNardo CD1, Rausch CR2, Benton C1, Kadia T1, Jain N1, Pemmaraju N1, Daver N1, Covert W2, Marx KR2, Mace M2, Jabbour E1, Cortes J1, Garcia-Manero G1, Ravandi F1, Bhalla KN1, Kantarjian H1, Konopleva M1.
American journal of hematology.Am J Hematol.2017 Dec 8. doi: 10.1002/ajh.25000. [Epub ahead of print]
PMID 29218851

Genetic classification of Aegilops columnaris Zhuk. (2n=4x=28, UcUcXcXc) chromosomes based on FISH analysis and substitution patterns in common wheat x Ae. columnaris introgressive lines.

Badaeva ED1,2, Ruban AS3, Shishkina AA4, Sibikeev SN5, Druzhin AE6, Surzhikov SA7, Dragovich AY8.
Genome.Genome.2017 Dec 7. doi: 10.1139/gen-2017-0186. [Epub ahead of print]
PMID 29216443

Japanese Journal

Adverse effects of inbreeding on colony foundation success in bumblebees, Bombus terrestris (Hymenoptera: Apidae)

Applied entomology and zoology 51(4), 521-526, 2016-11
NAID 40020985980

The Relationship between Mating System and Genetic Diversity in Diploid Sexual Populations of Cyrtomium falcatum in Japan

PLOS ONE 11(10), e0163683, 2016-10
NAID 120005893652

A Note on Natural Triploidy in a Japanese Brown Frog, Rana neba (Anura; Ranidae)

Current Herpetology 35(2), 128-131, 2016-08
NAID 120005848368

Relative DNA content in diploid, polyploid, and multiploid species of Paspalum (Poaceae) with relation to reproductive mode and taxonomy

Journal of plant research 129(4), 697-710, 2016-07
NAID 40020866933

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