Suidae
Temporal range: Oligocene–Recent
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Potamochoerus porcus |
Scientific classification |
Kingdom: |
Animalia |
Phylum: |
Chordata |
Class: |
Mammalia |
Order: |
Artiodactyla |
Suborder: |
Suina |
Family: |
Suidae
Gray, 1821 |
Genera |
Over 30 extinct genera, 6 extant,
see text.
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Suidae is the biological family to which pigs belong. In addition to numerous fossil species, up to sixteen extant species are currently recognized, classified into between four and eight genera. The family includes the domestic pig, Sus scrofa domesticus or Sus domesticus, in addition to numerous species of wild pig, such as the babirusa Babyrousa babyrussa and the warthog Phacochoerus aethiopicus. All suids are native to the Old World, ranging from Asia and its islands, to Europe, and Africa.
The earliest fossil suids date from the Oligocene epoch of Asia, and their descendants reached Europe during the Miocene.[1] Several fossil species are known, and show adaptations to a wide range of different diets, from strict herbivory to possible carrion-eating (in Tetraconodon).[2]
Contents
- 1 Physical characteristics
- 2 Behaviour and reproduction
- 3 Classification
- 4 See also
- 5 References
Physical characteristics[edit]
Suids belong to the order Artiodactyla, and are generally regarded as the living members of that order most similar to the ancestral form. Unlike most other members of the order, they have four toes on each foot, although they walk only on the middle two digits, with the others staying clear of the ground. They also have a simple stomach, rather than the more complex, ruminant, stomach found in most other Artiodactyl families.[3]
They are small to medium animals, varying in size from 58 to 66 cm (23 to 26 in) in length, and 6 to 9 kg (13 to 20 lb) in weight in the case of the Pygmy Hog, to 130–210 cm (4–7 ft) and 130–275 kg (300–600 lb) in the Giant Forest Hog. They have large heads and short necks, with relatively small eyes and prominent ears. Their heads have a distinctive snout, ending in a disc-shaped nose. Suids typically have a bristly coat, and a short tail ending in a tassle. The males possess a corkscrew-shaped penis, which fits into a similarly shaped groove in the female's cervix.
Suids have a well-developed sense of hearing, and are vocal animals, communicating with a series of grunts, squeals, and similar sounds. They also have an acute sense of smell. Many species are omnivorous, eating grass, leaves, roots, insects, worms, and even frogs or mice. Other species are more selective and purely herbivorous.[3]
Their teeth reflect their diet, and suids retain the upper incisors, which are lost in most other Artiodactyls. The canine teeth are enlarged to form prominent tusks, used for rooting in moist earth or undergrowth, and in fighting. They have only a short diastema. The number of teeth varies between species, but the general dental formula is: 1-3.1.2-4.33.1.2.3
Behaviour and reproduction[edit]
Wild boar feeding on carcass, Yala National Park.
Despite their apparently "primitive" anatomy, suids are intelligent and adaptable animals. Adult females (sows) and their young travel in a group (sounder; see List of animal names), while adult males (boars) are either solitary, or travel in small bachelor groups. Males generally are not territorial, and come into conflict only during the mating season.
Litter size varies between one and twelve, depending on the species. The mother prepares a grass nest or similar den, which the young leave after about ten days. Suids are weaned at around three months, and become sexually mature at 18 months. In practice, however, male suids are unlikely to gain access to sows in the wild until they have reached their full physical size, at around four years of age. In all species, the male is significantly larger than the female, and possesses more prominent tusks.[3]
Classification[edit]
Bearded Pig,
Sus barbatus
Chleuastochoerus fossil skull
The complete list of living species, and a partial list of extinct genera known from the fossil record, extinct taxa marked with a dagger "†",[2] are:
- Suidae
- Subfamily †Cainochoerinae
- Genus †Albanohyus
- Genus †Cainochoerus
- Subfamily †Hyotheriinae
- Genus †Aureliachoerus
- Genus †Chicochoerus
- Genus †Hyotherium
- Genus †Nguruwe (formerly placed in Kubanochoerinae)[4][5]
- Genus †Xenohyus
- Subfamily †Listriodontinae[4]
- Tribe †Kubanochoerini
- Genus †Kubanochoerus (junior synonyms Libycochoerus, Megalochoerus)
- Tribe †Listriodontini
- Genus †Eurolistriodon
- Genus †Listriodon (junior synonym Bunolistriodon)
- Tribe †Namachoerini
- Genus †Lopholistriodon
- Genus †Namachoerus
- Tribe incertae sedis
- Subfamily Suinae
- Tribe Babyrousini
- Genus Babyrousa (Pleistocene to recent)
- Species B. babyrussa Buru babirusa
- Species †B. bolabatuensis Bola Batu babirusa
- Species Babyrousa celebensis North Sulawesi babirusa
- Species Babyrousa togeanensis Togian babirusa
- Tribe †Hippohyini
- Genus †Hippohyus (Miocene to Pleistocene)
- Genus †Sinohyus (Miocene)
- Genus †Sivahyus (Miocene to Pliocene)
- Tribe Potamochoerini
- Genus †Celebochoerus (Pliocene to Pleistocene)
- Genus Hylochoerus (Pleistocene to recent)
- Species Hylochoerus meinertzhageni giant forest hog
- Genus †Kolpochoerus (Miocene to Pleistocene) (junior synonyms Ectopotamochoerus, Mesochoerus, Omochoerus, Promesochoerus)
- Genus Potamochoerus (Miocene to recent)
- Species P. larvatus bushpig
- Species P. porcus red river hog
- Genus †Propotamochoerus (Miocene to Pliocene)
- Tribe Suini
- Genus †Eumaiochoerus (Miocene)
- Genus †Hippopotamodon (Miocene to Pleistocene) (junior synonym Limnostonyx)
- Genus †Korynochoerus (Miocene to Pliocene)
- Genus †Microstonyx (Miocene)
- Genus Sus (Miocene to recent)
- Species S. ahoenobarbus Palawan bearded pig
- Species S. barbatus Bornean bearded pig
- Species S. bucculentus Heude's pig or Vietnamese warty pig
- Species S. cebifrons Visayan warty pig
- Species S. celebensis Celebes warty pig
- Species S. heureni Flores warty pig
- Species S. oliveri Mindoro warty pig
- Species S. philippensis Philippine warty pig
- Species S. scrofa (also called S. domesticus) domestic pig, wild boar
- Species S. verrucosus Java warty pig
- Species †S. strozzi
- Tribe Phacochoerini
- Genus †Metridiochoerus (Pliocene to Pleistocene)
- Genus Phacochoerus (Pliocene to recent)
- Species P. aethiopicus Cape, Somali or desert warthog
- Species P. africanus common warthog
- Genus †Potamochoeroides (Pliocene, possibly Pleistocene)
- Genus †Stylochoerus (Pleistocene)
- Tribe incertae sedis [6]
- Genus Porcula
- Species P. salvania pygmy hog
- Subfamily †Tetraconodontinae
- Genus †Conohyus
- Genus †Notochoerus
- Genus †Nyanzachoerus
- Genus †Parachleuastochoerus
- Genus †Sivachoerus
- Genus †Tetraconodon (Miocene, Myanmar) [7]
- Species †T. intermedius
- Species †T. malensis
- Species †T. minor
- Subfamily incertae sedis
- Genus †Chleuastochoerus[5]
- Genus †Hemichoerus
- Genus †Hyosus
- Genus †Kenyasus (formerly placed in Kubanochoerinae)[5]
- Genus †Schizochoerus[5]
- Genus †Sinapriculus[5]
See also[edit]
- Wild Pigs of the Philippines
References[edit]
- ^ Palmer, D., ed. (1999). The Marshall Illustrated Encyclopedia of Dinosaurs and Prehistoric Animals. London: Marshall Editions. p. 269. ISBN 1-84028-152-9.
- ^ a b Savage, RJG, & Long, MR (1986). Mammal Evolution: an illustrated guide. New York: Facts on File. pp. 212–213. ISBN 0-8160-1194-X.
- ^ a b c Cumming, David (1984). Macdonald, D., ed. The Encyclopedia of Mammals. New York: Facts on File. pp. 500–503. ISBN 0-87196-871-1.
- ^ a b Maeva, J.O. (2009). "The differentiation of bunodont Listriodontinae (Mammalia, Suidae) of Africa: new data from Kalodirr and Moruorot, Kenya". Zoological Journal of the Linnean Society 157 (3): 653–678. doi:10.1111/j.1096-3642.2008.00525.x.
- ^ a b c d e Maeva, J.O.; et al. (2010). "Phylogenetic relationships of the Suidae (Mammalia, Cetartiodactyla): new insights on the relationships within Suoidea". Zoologica Scripta 39 (4): 315–330. doi:10.1111/j.1463-6409.2010.00431.x.
- ^ Funk, S.M.; et al. (2007). "The pygmy hog is a unique genus: 19th century taxonomists got it right first time round". Molecular Phylogenetics and Evolution 45 (2): 427–436. doi:10.1016/j.ympev.2007.08.007. PMID 17905601.
- ^ Htike, T.; et al. (2005). "A revision of Tetraconodon (Mammalia, Artiodactyla, Suidae) from the Miocene of Myanmar and description of a new species". Paleontological Research 9 (3): 243–254. doi:10.2517/prpsj.9.243.
Extant Artiodactyla species
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- Kingdom: Animalia
- Phylum: Chordata
- Class: Mammalia
- Infraclass: Eutheria
- Superorder: Laurasiatheria
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Suborder Ruminantia
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Antilocapridae |
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Giraffidae |
Okapia
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Giraffa
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- Giraffe (G. camelopardalis)
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Moschidae |
Moschus
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- Himalayan musk deer (M. chrysogaster)
- Siberian musk deer (M. moschiferus)
- Dwarf musk deer (M. berezovskii)
- Black musk deer (M. fuscus)
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Tragulidae |
Hyemoschus
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- Water chevrotain (H. aquaticus)
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Moschiola
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- Indian spotted chevrotain (M. indica)
- Yellow-striped chevrotain (M. kathygre)
- Sri Lankan spotted chevrotain (M. meminna)
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Tragulus
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- Java mouse-deer (T. javanicus)
- Lesser mouse-deer (T. kanchil)
- Greater mouse-deer (T. napu)
- Philippine mouse-deer (T. nigricans)
- Vietnam mouse-deer (T. versicolor)
- Williamson's mouse-deer (T. williamsoni)
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Cervidae |
Large family listed below
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Bovidae |
Large family listed below
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Family Cervidae
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Cervinae |
Muntiacus
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- Indian muntjac (M. muntjak)
- Reeves's muntjac (M. reevesi)
- Hairy-fronted muntjac (M. crinifrons)
- Fea's muntjac (M. feae)
- Bornean yellow muntjac (M. atherodes)
- Roosevelt's muntjac (M. rooseveltorum)
- Gongshan muntjac (M. gongshanensis)
- Giant muntjac (M. vuquangensis)
- Truong Son muntjac (M. truongsonensis)
- Leaf muntjac (M. putaoensis)
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Elaphodus
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- Tufted deer (E. cephalophus)
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Dama
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- Fallow deer (D. dama)
- Persian fallow deer (D. mesopotamica)
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Axis
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Rucervus
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- Barasingha (R. duvaucelii)
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Panolia
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Elaphurus
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- Père David's deer (E. davidianus)
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Hyelaphus
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- Hog deer (H. porcinus)
- Calamian deer (H. calamianensis)
- Bawean deer (H. kuhlii)
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Rusa
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- Sambar deer (R. unicolor)
- Rusa deer (R. timorensis)
- Philippine sambar (R. mariannus)
- Philippine spotted deer (R. alfredi)
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Cervus
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- Red deer (C. elaphus)
- Elk (C. canadensis)
- Thorold's deer (C. albirostris)
- Sika deer (C. nippon)
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Capreolinae |
Alces
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Hydropotes
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Capreolus
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- Roe deer (C. capreolus)
- Siberian roe deer (C. pygargus)
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Rangifer
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Hippocamelus
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- Taruca (H. antisensis)
- South Andean deer (H. bisulcus)
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Mazama
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- Red brocket (M. americana)
- Small red brocket (M. bororo)
- Merida brocket (M. bricenii)
- Dwarf brocket (M. chunyi)
- Gray brocket (M. gouazoubira)
- Pygmy brocket (M. nana)
- Amazonian brown brocket (M. nemorivaga)
- Yucatan brown brocket (M. pandora)
- Little red brocket (M. rufina)
- Central American red brocket (M. temama)
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Ozotoceros
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- Pampas deer (O. bezoarticus)
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Blastocerus
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- Marsh deer (B. dichotomus)
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Pudu
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- Northern pudu (P. mephistophiles)
- Pudú (P. pudu)
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Odocoileus
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- White-tailed deer (O. virginianus)
- Mule deer (O. hemionus)
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Family Bovidae
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Cephalophinae |
Cephalophus
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- Abbott's duiker (C. spadix)
- Aders' duiker (C. adersi)
- Bay duiker (C. dorsalis)
- Black duiker (C. niger)
- Black-fronted duiker (C. nigrifrons)
- Blue duiker (C. monticola)
- Harvey's duiker (C. harveyi)
- Jentink's duiker (C. jentinki)
- Maxwell's duiker (C. maxwellii)
- Red forest duiker (C. natalensis)
- Ogilby's duiker (C. ogilbyi)
- Peters's duiker (C. callipygus)
- Red-flanked duiker (C. rufilatus)
- Ruwenzori duiker (C. rubidis)
- Weyns's duiker (C. weynsi)
- White-bellied duiker (C. leucogaster)
- White-legged duiker (C. crusalbum)
- Yellow-backed duiker (C. Sylvicultor)
- Zebra duiker (C. zebra)
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Sylvicapra
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- Common duiker (S. grimmia)
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Hippotraginae |
Hippotragus
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- Roan antelope (H. equinus)
- Sable antelope (H. niger)
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Oryx
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- East African oryx (O. beisa)
- Scimitar oryx (O. dammah)
- Gemsbok (O. gazella)
- Arabian oryx (O. leucoryx)
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Addax
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Reduncinae |
Kobus
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- Upemba lechwe (K. anselli)
- Waterbuck (K. ellipsiprymnus)
- Kob (K. kob)
- Lechwe (K. leche)
- Nile lechwe (K. megaceros)
- Puku (K. vardonii)
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Redunca
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- Southern reedbuck (R. arundinum)
- Mountain reedbuck (R. fulvorufula)
- Bohor reedbuck (R. redunca)
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Aepycerotinae |
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Peleinae |
Pelea
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- Grey rhebok (P. capreolus)
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Alcelaphinae |
Beatragus
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Damaliscus
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- Korrigum (D. korrigum)
- Common tsessebe (D. lunatus)
- Bontebok (D. pygargus)
- Bangweulu tsessebe (D. superstes)
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Alcelaphus
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- Hartebeest (A. buselaphus)
- Red hartebeest (A. caama)
- Lichtenstein's hartebeest (A. lichtensteinii)
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Connochaetes
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- Black wildebeest (C. gnou)
- Blue wildebeest (C. taurinus)
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Pantholopinae |
Pantholops
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- Tibetan antelope (P. hodgsonii)
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Caprinae |
Large subfamily listed below
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Bovinae |
Large subfamily listed below
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Antilopinae |
Large subfamily listed below
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Family Bovidae (subfamily Caprinae)
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Ammotragus
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- Barbary sheep (A. lervia)
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Budorcas
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Capra
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- Wild goat (C. aegagrus)
- West Caucasian tur (C. caucasia)
- East Caucasian tur (C. cylindricornis)
- Markhor (C. falconeri)
- Alpine ibex (C. ibex)
- Nubian ibex (C. nubiana)
- Spanish ibex (C. pyrenaica)
- Siberian ibex (C. sibirica)
- Walia ibex (C. walie)
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Hemitragus
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- Nilgiri tahr (H. hylocrius)
- Arabian tahr (H. jayakari)
- Himalayan tahr (H. jemlahicus)
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Naemorhedus
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- Red goral (N. baileyi)
- Japanese serow (N. crispus)
- Long-tailed goral (N. caudatus)
- Himalayan goral (N. goral)
- Mainland serow (N. milneedwardsii)
- Taiwan serow (N. swinhoei)
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Oreamnos
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- Mountain goat (O. americanus)
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Ovibos
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Ovis
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- Argali (O. ammon)
- Domestic sheep (O. aries)
- Bighorn sheep (O. canadensis)
- Dall sheep (O. dalli)
- Mouflon (O. musimon)
- Snow sheep (O. nivicola)
- Urial (O. orientalis)
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Pseudois
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- Bharal (P. nayaur)
- Dwarf blue sheep (P. schaeferi)
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Rupicapra
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- Pyrenean chamois (R. pyrenaica)
- Chamois (R. rupicapra)
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Family Bovidae (subfamily Bovinae)
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Boselaphini |
Tetracerus
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- Four-horned antelope (T. quadricornis)
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Boselaphus
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Bovini |
Bubalus
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- Water buffalo (B. bubalus)
- Wild Water Buffalo (B. arnee)
- Lowland anoa (B. depressicornis)
- Mountain anoa (B. quarlesi)
- Tamaraw (B. mindorensis)
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Bos
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- Banteng (B. javanicus)
- Gaur (B. gaurus)
- Yak (B. mutus)
- Cattle (B. taurus)
- Kouprey (B. sauveli)
- Zebu (B. indicus)
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Pseudonovibos
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Pseudoryx
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Syncerus
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- African buffalo (S. caffer)
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Bison
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- American bison (B. bison)
- Wisent (B. bonasus)
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Strepsicerotini |
Tragelaphus
(including kudus)
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- Sitatunga (T. spekeii)
- Nyala (T. angasii)
- Bushbuck (T. scriptus)
- Mountain nyala (T. buxtoni)
- Lesser kudu (T. imberbis)
- Greater kudu (T. strepsiceros)
- Bongo (T. eurycerus)
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Taurotragus
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- Common eland (T. oryx)
- Giant eland (T. derbianus)
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Family Bovidae (subfamily Antilopinae)
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Antilopini |
Ammodorcas
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Antidorcas
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- Springbok (A. marsupialis)
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Antilope
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- Blackbuck (A. cervicapra)
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Gazella
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- Mountain gazelle (G. gazella)
- Neumann's gazelle (G. erlangeri)
- Speke's gazelle (G. spekei)
- Dorcas gazelle (G. dorcas)
- Saudi gazelle (G. saudiya)
- Chinkara (G. bennettii)
- Thomson's gazelle (G. thomsonii)
- Red-fronted gazelle (G. rufifrons)
- Dama gazelle (G. dama)
- Grant's gazelle (G. granti)
- Soemmerring's gazelle (G. soemmerringii)
- Cuvier's gazelle (G. cuvieri)
- Rhim gazelle (G. leptoceros)
- Goitered gazelle (G. subgutturosa)
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Litocranius
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Procapra
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- Mongolian gazelle (P. gutturosa)
- Goa (P. picticaudata)
- Przewalski's gazelle (P. przewalskii)
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Saigini |
Pantholops
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- Tibetan antelope (P. hodgsonii)
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Saiga
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- Saiga antelope (S. tatarica)
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Neotragini |
Dorcatragus
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Madoqua
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- Günther's dik-dik (M. guentheri)
- Kirk's dik-dik (M. kirkii)
- Silver dik-dik (M. piacentinii)
- Salt's dik-dik (M. saltiana)
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Neotragus
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- Bates's pygmy antelope (N. batesi)
- Suni (N. moschatus)
- Royal antelope (N. pygmaeus)
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Oreotragus
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- Klipspringer (O. oreotragus)
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Ourebia
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Raphicerus
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- Steenbok (R. campestris)
- Cape grysbok (R. melanotis)
- Sharpe's grysbok (R. sharpei)
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Suborder Suina
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Suidae |
Babyrousa
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- Buru babirusa (B. babyrussa)
- North Sulawesi babirusa (B. celebensis)
- Togian babirusa (B. togeanensis)
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Hylochoerus
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- Giant forest hog (H. meinertzhageni)
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Phacochoerus
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- Desert warthog (P. aethiopicus)
- Warthog (P. africanus)
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Porcula
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Potamochoerus
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- Bushpig (P. larvatus)
- Red River Hog (P. porcus)
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Sus
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- Palawan bearded pig (S. ahoenobarbus)
- Bornean bearded pig (S. barbatus)
- Indo-chinese warty pig (S. bucculentus)
- Visayan warty pig (S. cebifrons)
- Celebes warty pig (S. celebensis)
- Flores warty pig (S. heureni)
- Oliver's warty pig (S. oliveri)
- Philippine warty pig (S. philippensis)
- Wild boar (S. scrofa)
- Timor warty pig (S. timoriensis)
- Javan warty pig (S. verrucosus)
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Tayassuidae |
Tayassu
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- White-lipped peccary (T. pecari)
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Catagonus
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- Chacoan peccary (C. wagneri)
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Pecari
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- Collared peccary (P. tajacu)
- Giant peccary (P. maximus)
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Suborder Tylopoda
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Camelidae
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Lama
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- Llama (L. glama)
- Guanaco (L. guanicoe)
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Vicugna
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- Vicuña (V. vicugna)
- Alpaca (V. pacos)
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Camelus
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- Dromedary (C. dromedarius)
- Bactrian camel (C. bactrianus)
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Cetartiodactyla (unranked clade, higher than Artiodactyla)
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Hippopotamidae
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Hippopotamus
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- Hippopotamus (H. amphibius)
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Choeropsis
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- Pygmy hippopotamus (C. liberiensis)
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