An infant born to an alcoholic mother had microcephaly and cardiac abnormalities and died despite resuscitative efforts.
vi.
持ちこたえる。耐える
圧迫する(on, against)。(支持物に)もたれかかる(on)
効き目がある(on)。関係/影響がある(on)
押し進む。(ある方向を)とる。向く、曲がる
WordNet
move while holding up or supporting; "Bear gifts"; "bear a heavy load"; "bear news"; "bearing orders"
massive plantigrade carnivorous or omnivorous mammals with long shaggy coats and strong claws
an investor with a pessimistic market outlook; an investor who expects prices to fall and so sells now in order to buy later at a lower price
have rightfully; of rights, titles, and offices; "She bears the title of Duchess"; "He held the governorship for almost a decade" (同)hold
take on as ones own the expenses or debts of another person; "Ill accept the charges"; "She agreed to bear the responsibility" (同)take_over, accept, assume
bring forth, "The apple tree bore delicious apples this year"; "The unidentified plant bore gorgeous flowers" (同)turn_out
have; "bear a resemblance"; "bear a signature"
remain inactive or immobile; "standing water"
a defensive effort; "the army made a final stand at the Rhone"
a small table for holding articles of various kinds; "a bedside stand"
a stop made by a touring musical or theatrical group to give a performance; "a one-night stand"
tiered seats consisting of a structure (often made of wood) where people can sit to watch an event (game or parade)
a growth of similar plants (usually trees) in a particular area; "they cut down a stand of trees"
the position where a thing or person stands
hold ones ground; maintain a position; be steadfast or upright; "I am standing my ground and wont give in!" (同)remain firm
be standing; be upright; "We had to stand for the entire performance!" (同)stand_up
put into an upright position; "Can you stand the bookshelf up?" (同)stand_up, place upright
be available for stud services; "male domestic animals such as stallions serve selected females"
be in effect; be or remain in force; "The law stands!"
be in some specified state or condition; "I stand corrected"
be tall; have a height of; copula; "She stands 6 feet tall"
have or maintain a position or stand on an issue; "Where do you stand on the War?"
occupy a place or location, also metaphorically; "We stand on common ground"
have left; "I have two years left"; "I dont have any money left"; "They have two more years before they retire"
undergo; "The stocks had a fast run-up" (同)experience
suffer from; be ill with; "She has arthritis"
have as a feature; "This restaurant features the most famous chefs in France" (同)feature
achieve a point or goal; "Nicklaus had a 70"; "The Brazilian team got 4 goals"; "She made 29 points that day" (同)get, make
have or possess, either in a concrete or an abstract sense; "She has $1,000 in the bank"; "He has got two beautiful daughters"; "She holds a Masters degree from Harvard" (同)have got, hold
be confronted with; "What do we have here?"; "Now we have a fine mess"
have a personal or business relationship with someone; "have a postdoc"; "have an assistant"; "have a lover"
recognize and respect (rights and beliefs of others); "We must tolerate the religions of others"
have a tolerance for a poison or strong drug or pathogen or environmental condition; "The patient does not tolerate the anti-inflammatory drugs we gave him"
relating to or connected with or occurring at the time of childbirth or shortly following, or to the woman who has just given birth
only the giant panda: in some classifications considered a genus of the separate family Ailuropodidae (同)genus Ailuropoda
overcome; "overbear criticism, protest, or arguments"
bear too much
the attendant (usually an officer of a noblemans household) whose duty is to fill and serve cups of wine
large shaggy-haired toothless anteater with long tongue and powerful claws; of South America (同)giant anteater, great anteater, tamanoir, Myrmecophaga jubata
North American astilbe with panicles of creamy white flowers (同)Astilbe biternata
to remain unmolested, undisturbed, or uninterrupted -- used only in infinitive form; "let her be"
work in a specific place, with a specific subject, or in a specific function; "He is a herpetologist"; "She is our resident philosopher" (同)follow
have life, be alive; "Our great leader is no more"; "My grandfather lived until the end of war" (同)live
be identical to; be someone or something; "The president of the company is John Smith"; "This is my house"
happen, occur, take place; "I lost my wallet; this was during the visit to my parents house"; "There were two hundred people at his funeral"; "There was a lot of noise in the kitchen"
have the quality of being; (copula, used with an adjective or a predicate noun); "John is rich"; "This is not a good answer"
occupy a certain position or area; be somewhere; "Where is my umbrella?" "The toolshed is in the back"; "What is behind this behavior?"
spend or use time; "I may be an hour"
relevant relation or interconnection; "those issues have no bearing on our situation"
(of a structural member) withstanding a weight or strain
《現在完了》《have(has)+過去分詞》・《現在における動作の完了・結果》…『した』,…『してしまった』・《現在までの経験》…『したことがある』・《現在までの継続》(ずっと)…『している』,…『してきた』・《時・条件の副詞節で未来完了に代えて》…してしまう・《過去完了》《had+過去分詞》・《未来完了》《will(shall) have +過去分詞》・《未来のある時における動作の完了・結果》…『して』[『しまって』]『いるだろう』・《未来のある時までの経験》…『したことになる』・《未来のある時までの継続》…『していることになる』・《完了形不定詞》・《述語動詞の表す時より前に事柄を表す》・《期待・意図・願望・予定などを表す動詞の過去形に付いて実現されなかった事柄を表す》・《完了形分詞または動名詞》《述語動詞の表す時までに完了した事柄またはその結果を表す》
This article is about the mammalian family. For the koala and extinct bear-like marsupials, see Phascolarctidae. For other uses, see Bear (disambiguation).
"Ursine" redirects here. For the village, see Ursine, Nevada.
Bears
Temporal range: 38–0Ma
PreЄ
Є
O
S
D
C
P
T
J
K
Pg
N
Late Eocene – Recent
Brown bear, Ursus arctos in Norway
Scientific classification
Kingdom:
Animalia
Phylum:
Chordata
Class:
Mammalia
Order:
Carnivora
Suborder:
Caniformia
Family:
Ursidae
G. Fischer de Waldheim, 1817
Genera
Ailuropoda
Helarctos
Melursus
Ursus
Tremarctos
Agriarctos †
Agriotherium †
Amphicticeps †
Amphicynodon †
Arctotherium †
Arctodus †
Indarctos †
Kolponomos †
Parictis †
Plionarctos †
Ursavus †
Bears are mammals of the family Ursidae. Bears are classified as caniforms, or doglike carnivorans, with the pinnipeds being their closest living relatives. Although only eight species of bears are extant, they are widespread, appearing in a wide variety of habitats throughout the Northern Hemisphere and partially in the Southern Hemisphere. Bears are found on the continents of North America, South America, Europe, and Asia.
Common characteristics of modern bears include large bodies with stocky legs, long snouts, shaggy hair, plantigrade paws with five nonretractile claws, and short tails. While the polar bear is mostly carnivorous and the giant panda feeds almost entirely on bamboo, the remaining six species are omnivorous, with varied diets.
With the exceptions of courting individuals and mothers with their young, bears are typically solitary animals. They are generally diurnal, but may be active during the night (nocturnal) or twilight (crepuscular), particularly around humans. Bears are aided by an excellent sense of smell, and despite their heavy build and awkward gait, they can run quickly and are adept climbers and swimmers. In autumn, some bear species forage large amounts of fermented fruits, which affects their behaviour.[1] Bears use shelters, such as caves and burrows, as their dens; most species occupy their dens during the winter for a long period of sleep similar to hibernation.
Bears have been hunted since prehistoric times for their meat and fur. To this day, they play a prominent role in the arts, mythology, and other cultural aspects of various human societies. In modern times, the bears' existence has been pressured through the encroachment on their habitats and the illegal trade of bears and bear parts, including the Asian bile bear market. The IUCN lists six bear species as vulnerable or endangered, and even least concern species, such as the brown bear, are at risk of extirpation in certain countries. The poaching and international trade of these most threatened populations are prohibited, but still ongoing.
Contents
1Etymology
2Evolutionary history
2.1Fossil bears
2.2Taxonomic revisions of living bear species
3Classification
4Biology
4.1Morphology
4.1.1Dentition
4.2Distribution and habitat
4.3Behaviour
4.4Vocalizations
4.5Diet and interspecific interactions
4.6Breeding
4.7Winter dormancy
5Relationship with humans
5.1Bear hunt
6Culture
6.1Names
6.2Myth and legend
6.3Symbolic use
6.4Figures of speech
6.5Teddy bears
7Organizations regarding bears
8See also
9References
10Further reading
11External links
Etymology
The English word "bear" comes from Old English bera and belongs to a family of names for the bear in Germanic languages that originate from an adjective meaning "brown".[2] In Scandinavia, the word for bear is björn (or bjørn), and is a relatively common given name for males. The use of this name is ancient and has been found mentioned in several runestone inscriptions.[3]
The reconstructed Proto-Indo-European name of the bear is *h₂ŕ̥tḱos, whence Sanskrit r̥kṣa, Avestan arša, Greek ἄρκτος (arktos), Latin ursus, Welsh arth (whence perhaps "Arthur"), Albanian ari, Armenian արջ (arj). Also compared is Hittite ḫartagga-, the name of a monster or predator.[2] In the binomial name of the brown bear, Ursus arctos, Linnaeus simply combined the Latin and Greek names.
The Proto-Indo-European (PIE) word for bear, *h₂ŕ̥tḱos seems to have been subject to taboo deformation or replacement in some languages (as was the word for wolf, wlkwos), resulting in the use of numerous unrelated words with meanings like "brown one" (English bruin) and "honey-eater" (Slavic medved).[4] Thus, some Indo-European language groups do not share the same PIE root. The theory of the bear taboo is taught to almost all starting students of Indo-European and historical linguistics; the putative original PIE word for bear is itself descriptive, because a cognate word in Sanskrit is rakṣas, meaning "harm, injury".[5]
Evolutionary history
The family Ursidae is one of 9 families in the suborder Caniformia, or "doglike" carnivores, within the order Carnivora. Bears' closest living relatives are the pinnipeds, canids, and musteloids.[6]
The following synapomorphic (derived) traits set bears apart from related families:
presence of an alisphenoid canal
paroccipital processes that are large and not fused to the auditory bullae
auditory bullae are not enlarged
lacrimal bone is vestigial
cheek teeth are bunodont and hence indicative of a broad, hypocarnivorous (not strictly meat-eating) diet (although hypercarnivorous (strictly meat-eating) taxa are known from the fossil record)[7]
carnassials are flattened
Additionally, members of this family possess posteriorly oriented M2 postprotocrista molars, elongated m2 molars, and a reduction of the premolars.
Modern bears comprise eight species in three subfamilies: Ailuropodinae (monotypic with the giant panda), Tremarctinae (monotypic with the spectacled bear), and Ursinae (containing six species divided into one to three genera, depending upon authority).
Fossil bears
The earliest members of Ursidae belong to the extinct subfamily Amphicynodontinae, including Parictis (late Eocene to early middle Miocene, 38–18 Mya) and the slightly younger Allocyon (early Oligocene, 34–30 Mya), both from North America. These animals looked very different from today's bears, being small and raccoon-like in overall appearance, and diets perhaps more similar to that of a badger. Parictis does not appear in Eurasia and Africa until the Miocene.[8] It is unclear whether late-Eocene ursids were also present in Eurasia, although faunal exchange across the Bering land bridge may have been possible during a major sea level low stand as early as the late Eocene (about 37 Mya) and continuing into the early Oligocene.[9] European genera morphologically are very similar to Allocyon, and also the much younger American Kolponomos (about 18 Mya), are known from the Oligocene, including Amphicticeps and Amphicynodon.
Plithocyon armagnacensis skull
The raccoon-sized, dog-like Cephalogale is the oldest-known member of the subfamily Hemicyoninae, which first appeared during the middle Oligocene in Eurasia about 30 Mya ago. The subfamily also includes the younger genera Phoberocyon (20–15 Mya), and Plithocyon (15–7 Mya).
A Cephalogale-like species gave rise to the genus Ursavus during the early Oligocene (30–28 Mya); this genus proliferated into many species in Asia and is ancestral to all living bears. Species of Ursavus subsequently entered North America, together with Amphicynodon and Cephalogale, during the early Miocene (21–18 Mya).
Members of the living lineages of bears diverged from Ursavus around 20 Mya ago, likely via the species Ursavus elmensis. Based on genetic and morphological data, the Ailuropodinae (pandas) were the first to diverge from other living bears about 19 Mya ago, although no fossils of this group have been found before about 5 Mya.[10]
The New World short-faced bears (Tremarctinae) differentiated from Ursinae following a dispersal event into North America during the mid Miocene (about 13 Mya).[10] They invaded South America (~1 Ma) following formation of the Isthmus of Panama.[11] Their earliest fossil representative is Plionarctos in North America (~ 10–2 Ma). This genus is probably the direct ancestor to the North American short-faced bears (genus Arctodus), the South American short-faced bears (Arctotherium), and the spectacled bears, Tremarctos, represented by both an extinct North American species (T. floridanus), and the lone surviving representative of the Tremarctinae, the South American spectacled bear (T. ornatus).
Fossil of cave bear (Ursus spelaeus)
The subfamily Ursinae experienced a dramatic proliferation of taxa about 5.3–4.5 Mya ago coincident with major environmental changes, with the first members of the genus Ursus also appearing around this time.[10] The sloth bear is a modern survivor of one of the earliest lineages to diverge during this radiation event (5.3 Mya); it took on its peculiar morphology, related to its diet of termites and ants, no later than by the early Pleistocene. By 3–4 Mya ago, the species Ursus minimus appears in the fossil record of Europe; apart from its size, it was nearly identical to today's Asiatic black bear. It is likely ancestral to all bears within Ursinae, perhaps aside from the sloth bear. Two lineages evolved from U. minimus: the black bears (including the sun bear, the Asiatic black bear, and the American black bear); and the brown bears (which includes the polar bear). Modern brown bears evolved from U. minimus via Ursus etruscus, which itself is ancestral to both the extinct Pleistocene cave bear and today's brown and polar bears. Species of Ursinae have migrated repeatedly into North America from Eurasia as early as 4 Mya during the early Pliocene.[12]
The fossil record of bears is exceptionally good. Direct ancestor-descendent relationships between individual species are often fairly well established, with sufficient intermediate forms known to make the precise cut-off between an ancestral and its daughter species subjective.[13]
Other extinct bear genera include Agriarctos, Indarctos, and Agriotherium (sometimes placed within hemicyonids).
Taxonomic revisions of living bear species
The giant panda's taxonomy (subfamily Ailuropodinae) has long been debated. Its original classification by Armand David in 1869 was within the bear genus Ursus, but in 1870, it was reclassified by Alphonse Milne-Edwards to the raccoon family.[14] In recent studies, the majority of DNA analyses suggest the giant panda has a much closer relationship to other bears and should be considered a member of the family Ursidae.[15] Estimates of divergence dates place the giant panda as the most ancient offshoot among living taxa within Ursidae, having split from other bears 17.9 to 22.1 Mya.[10] The red panda was included within Ursidae in the past. However, more recent research does not support such a conclusion, and instead places it in its own family Ailuridae, in superfamily Musteloidea along with Mustelidae, Procyonidae, and Mephitidae.[16][17][18] Multiple similarities between the two pandas, including the presence of false thumbs, are thus thought to represent an example of convergent evolution for feeding primarily on bamboo.
Unlike their neighbors elsewhere, the brown bears of Alaska's ABC Islands evidently are more closely related to polar bears than to other brown bears in the world. Researchers Gerald Shields and Sandra Talbot of the University of Alaska Fairbanks Institute of Arctic Biology studied the DNA of several samples of the species and found their DNA is different from that of other brown bears. The discovery has shown, while all other brown bears share a brown bear as their closest relative, those of Alaska's ABC Islands differ and share their closest relation with the polar bear.[19] Also, the very rare Tibetan blue bear is a type of brown bear. This animal has never been photographed.[citation needed]
Koalas are often referred to as bears due to their appearance; they are not bears, however, but are marsupials.
Classification
Brown bear (Ursus arctos) at the Moscow Zoo
Asian black bear Ursus thibetanus, at the Wrocław Zoo, Poland
Sun bear (Helarctos malayanus) at the Columbus Zoo and Aquarium
Giant panda (Ailuropoda melanoleuca) at the Smithsonian National Zoological Park, Washington DC
Family Ursidae
Subfamily Ailuropodinae
† Ailurarctos
† Ailurarctos lufengensis
† Ailurarctos yuanmouenensis
Ailuropoda (pandas)
† Ailuropoda baconi
† Ailuropoda fovealis
Ailuropoda melanoleuca, giant panda
Ailuropoda melanoleuca melanoleuca, giant panda
Ailuropoda melanoleuca qinlingensis, Qinling panda
† Ailuropoda microta
† Ailuropoda wulingshanensis
Subfamily Tremarctinae
Tremarctos (Spectacled bears)
Tremarctos ornatus, spectacled bear
† Tremarctos floridanus
† Arctodus
† Arctodus simus
† Arctodus pristinus
† Arctotherium
† Arctotherium angustidens
† Arctotherium bonariense
† Arctotherium brasilense
† Arctotherium latidens
† Arctotherium tarijense
† Arctotherium vetustum
† Arctotherium wingei
† Plionarctos
† Plionarctos edensis
† Plionarctos harroldorum
Subfamily Ursinae
† Agriotherium
† Agriotherium inexpetans
† Agriotherium schneideri
† Agriotherium sivalensis
Helarctos
Helarctos malayanus, sun bear
Helarctos malayanus malayanus
Helarctos malayanus euryspilus, Borneo sun bear
† Indarctos
† Indarctos anthraciti
† Indarctos arctoides
† Indarctos atticus
† Indarctos nevadensis
† Indarctos oregonensis
† Indarctos salmontanus
† Indarctos vireti
† Indarctos zdanskyi
Melursus
Melursus ursinus, sloth bear
Melursus ursinus inornatus, Sri Lankan sloth bear
Melursus ursinus ursinus, Indian sloth bear
† Ursavus
† Ursavus brevirhinus
† Ursavus depereti
† Ursavus elmensis
† Ursavus pawniensis
† Ursavus primaevus
Ursus
† Ursus abstrusus
Ursus americanus, American black bear
Ursus americanus altifrontalis, Olympic black bear
Ursus americanus amblyceps, New Mexico black bear
Ursus americanus americanus, Eastern black bear
Ursus americanus californiensis, California black bear
Ursus americanus carlottae, Haida Gwaii black bear or Queen Charlotte black bear
Ursus americanus cinnamomum, cinnamon bear
Ursus americanus emmonsii, Glacier bear
Ursus americanus eremicus, Mexican black bear
Ursus americanus floridanus, Florida black bear
Ursus americanus hamiltoni, Newfoundland black bear
Ursus americanus kermodei, Kermode bear or spirit bear
Ursus americanus luteolus, Louisiana black bear
Ursus americanus machetes, West Mexico black bear
Ursus americanus perniger, Kenai black bear
Ursus americanus pugnax, Dall black bear
Ursus americanus vancouveri, Vancouver Island black bear
Ursus arctos, brown bear
Ursus arctos arctos, Eurasian brown bear
Ursus arctos alascensis
Ursus arctos beringianus, Kamchatka brown bear or Far Eastern brown bear
† Ursus arctos californicus, California golden bear
† Ursus arctos crowtheri, Atlas bear
† Ursus arctos dalli
Ursus arctos gobiensis, Gobi bear (very rare)
Ursus arctos horribilis, grizzly bear
Ursus arctos isabellinus, Himalayan brown bear or Himalayan red bear
Ursus arctos lasiotus, Ussuri brown bear or black grizzly
Ursus arctos middendorffi, Kodiak bear
† Ursus arctos nelsoni, Mexican grizzly bear
Ursus arctos piscator, Bergman's bear (extinct?)
Ursus arctos pruinosus, Tibetan blue bear, Tibetan bear, or Himalayan blue bear
Ursus arctos sitkensis
Ursus arctos syriacus Syrian (brown) bear
† Ursus deningeri
† Ursus etruscus
† Ursus inopinatus, MacFarlane's bear (cryptid; if an authentic species, extinct)
† Ursus minimus
Ursus maritimus, polar bear
Ursus maritimus maritimus
† Ursus maritimus tyrannus
† Ursus rossicus
† Ursus sackdillingensis
† Ursus spelaeus, cave bear
Ursus thibetanus, Asian black bear
Ursus thibetanus formosanus, Formosan black bear
Ursus thibetanus gedrosianus
Ursus thibetanus japonicus
Ursus thibetanus laniger
Ursus thibetanus mupinensis
Ursus thibetanus thibetanus
Ursus thibetanus ussuricus
† Kolponomos
† Kolponomos clallamensis
† Kolponomos newportensis
The genera Melursus and Helarctos are sometimes also included in Ursus. The Asiatic black bear and the polar bear used to be placed in their own genera, Selenarctos and Thalarctos; these names have since been reduced in rank to subgeneric rank.
A number of hybrids have been bred between American black, brown, and polar bears.
Biology
Morphology
Despite being quadrupeds, bears can stand and sit similarly to humans.
Unlike most other Carnivora, bears have plantigrade feet
Bears are generally bulky and robust animals with relatively short legs. They are sexually dimorphic with regard to size, with the males being larger. Larger species tend to show increased levels of sexual dimorphism in comparison to smaller species, and where a species varies in size across its distribution, individuals from larger-sized areas tend also to vary more. Bears are the most massive terrestrial members of the order Carnivora. Some exceptional polar bears and Kodiak bears (a brown bear subspecies) have been weighed at over 750 kg (1,700 lb). As to which species is the largest depends on whether the assessment is based on which species has the largest individuals (brown bears) or on the largest average size (polar bears), as some races of brown bears are much smaller than polar bears. Adult male Kodiak bears average 480 to 533 kg (1,100 to 1,180 lb) compared to an average of 386 to 408 kg (850 to 900 lb) in adult male polar bears, per the Guinness Book of World Records.[20] The smallest bears are the sun bears of Asia, which weigh an average of 65 kg (140 lb) for the males and 45 kg (99 lb) for the females, though the smallest mature females can weigh only 20 kg (44 lb).[21][22] All "medium"-sized bear species (which include the other five extant species) are around the same average weight, with males averaging around 100 to 120 kg (220 to 260 lb) and females averaging around 60 to 85 kg (130 to 190 lb), although it is not uncommon for male American black bears to considerably exceed "average" weights.[23] Head-and-body length can range from 120 cm (47 in) in sun bears to 300 cm (120 in) in large polar and brown bears and shoulder height can range from 60 cm (24 in) to over 160 cm (63 in) in the same species, respectively. The tails of bears are often considered a vestigal feature and can range from 3 to 22 cm (1.2 to 8.7 in).[22][23]
Unlike most other land carnivorans, bears are plantigrade. They distribute their weight toward the hind feet, which makes them look lumbering when they walk. They are still quite fast, with the brown bear reaching 30 mph (48 km/h), although they are still slower than felines and canines. Bears can stand on their hind feet and sit up straight with remarkable balance. Bears' nonretractable claws are used for digging, climbing, tearing, and catching prey. Their ears are rounded.
Bears have an excellent sense of smell, better than the dogs (Canidae), or possibly any other mammal. This sense of smell is used for signalling between bears (either to warn off rivals or detect mates) and for finding food. Smell is the principal sense used by bears to find most of their food.[21]
Dentition
Unlike most other members of the Carnivora, bears have relatively undeveloped carnassial teeth, and their teeth are adapted for a diet that includes a significant amount of vegetable matter. The canine teeth are large, and the molar teeth flat and crushing. Considerable variation occurs in dental formula even within a given species. This may indicate bears are still in the process of evolving from carnivorous to predominantly herbivorous diets. Polar bears appear to have secondarily re-evolved fully functional carnassials, as their diets have switched back towards carnivory.[24] The dental formula for living bears is:
Distribution and habitat
See also: List of carnivorans by population
Bears are primarily found in the Northern Hemisphere, and with one exception, only in Asia, North America and Europe. A single species, the spectacled bear, is native to the Andean region of South America. The Atlas bear, a subspecies of the brown bear, was the only bear native to Africa. It was distributed in North Africa from Morocco to Libya, but has been extinct since around the 1870s. The most widespread species is the brown bear, which occurs from Western Europe eastwards through Asia to the western areas of North America. The American black bear is restricted to North America, and the polar bear is restricted to the Arctic Sea. All the remaining species are Asian.[21]
With the exception of the polar bear, bears are mostly forest species. Some species, particularly the brown bear, may inhabit or seasonally use other areas, such as alpine scrub or tundra.
Behaviour
While many people think bears are nocturnal, they are, in fact, generally diurnal, active for the most part during the day. The belief they are nocturnal apparently comes from the habits of bears that live near humans, which engage in some nocturnal activities, such as raiding trash cans or crops while avoiding humans. The sloth bear of Asia is the most nocturnal of the bears, but this varies by individual, and females with cubs are often diurnal to avoid competition with males and nocturnal predators.[21] Bears are overwhelmingly solitary and are considered to be the most asocial of all the Carnivora. Liaisons between breeding bears are brief, and the only times bears are encountered in small groups are mothers with young or occasional seasonal bounties of rich food (such as salmon runs).[21]
Vocalizations
Bears produce a variety of vocalizations such as:
Moaning is produced mostly as mild warnings to potential threats or in fear,
Barking is produced during times of alarm, excitement or to give away the animal's position.
Huffing is made during courtship or between mother and cubs to warn of danger.
Growling is produced as strong warnings to potential threats or in anger.
Roaring is used much for the same reasons as growls and also to proclaim territory and for intimidation.
Diet and interspecific interactions
Asian black bear feeding on berries
Brown bears make use of infrequent but predictable salmon runs in order to feed
Their carnivorous reputation not withstanding, most bears have adopted diets of more plant than animal matter and are completely opportunistic omnivores. Some bears will climb trees to obtain mast (edible vegatative or reproductive parts such as acorns); smaller species that are more able to climb include a greater amount of this in their diets.[25] Such masts can be very important to the diets of these species, and mast failures may result in long-range movements by bears looking for alternative food sources.[26] One exception is the polar bear, which has adopted a diet mainly of marine mammals to survive in the Arctic. The other exception is the giant panda, which has adopted a diet mainly of bamboo. Stable isotope analysis of the extinct giant short-faced bear (Arctodus simus) shows it was also an exclusive meat-eater, probably a scavenger.[27] The sloth bear, though not as specialized as the previous two species, has lost several front teeth usually seen in bears, and developed a long, suctioning tongue to feed on the ants, termites, and other burrowing insects they favour. At certain times of the year, these insects can make up 90% of their diets.[28] All bears will feed on any food source that becomes available, the nature of which varies seasonally. A study of Asiatic black bears in Taiwan found they would consume large numbers of acorns when they were most common, and switch to ungulates at other times of the year.[29]
When taking warm-blooded animals, bears will typically take small or young animals, as they are easier to catch. However, both species of black bears and the brown bear can sometimes take large prey, such as ungulates.[29][30] Often, bears will feed on other large animals when they encounter a carcass, whether or not the carcass is claimed by, or is the kill of, another predator. This competition is the main source of interspecies conflict. Bears are typically the apex predators in their ranges due to their size and power, and can defend a carcass against nearly all comers. Mother bears also can usually defend their cubs against other predators. The tiger is the only predator known to regularly prey on adult bears, including sloth bears, Asiatic black bears, giant pandas, sun bears and small brown bears.
Breeding
Bear cubs, like this American black bear, are sometimes killed by males.
The age at which bears reach sexual maturity is highly variable, both between and within species. Sexual maturity is dependent on body condition, which is in turn dependent upon the food supply available to the growing individual. The females of smaller species may have young in as little as two years, whereas the larger species may not rear young until they are four or even 9 years old. First breeding may be even later in males, where competition for mates may leave younger males without access to females.[21]
American black bears mating
The bear's courtship period is very brief. Bears in northern climates reproduce seasonally, usually after a period of inactivity similar to hibernation, although tropical species breed all year round. Cubs are born toothless, blind, and bald. The cubs of brown bears, usually born in litters of one to three, will typically stay with the mother for two full seasons. They feed on their mother's milk through the duration of their relationship with their mother, although as the cubs continue to grow, nursing becomes less frequent and cubs learn to begin hunting with the mother. They will remain with the mother for about three years, until she enters the next cycle of estrus and drives the cubs off. Bears will reach sexual maturity in five to seven years. Male bears, especially polar and brown bears, will kill and sometimes devour cubs born to another father to induce a female to breed again. Female bears are often successful in driving off males in protection of their cubs, despite being rather smaller.
Winter dormancy
Polar bear mother nursing her cub
Polar bear at Wapusk National Park, Canada
Many bears of northern regions are assumed[by whom?] to hibernate in the winter. While many bear species do go into a physiological state often colloquially called "hibernation" or "winter sleep", it is not true hibernation.[citation needed] In true hibernators, body temperatures drop to near ambient and heart rates slow drastically, but the animals periodically rouse themselves to urinate or defecate and to eat from stored food. The body temperature of bears, on the other hand, drops only a few degrees from normal and heart rate slows only slightly. They normally do not wake during this "hibernation", so do not eat, drink, urinate, or defecate the entire period. Higher body heat and being easily roused may be adaptations, because females give birth to their cubs during this winter sleep.
Relationship with humans
See also: Bear danger
Some species, such as the polar bear, American black bear, sloth bear, and brown bear, are dangerous to humans, especially in areas where they have become used to people. All bears are physically powerful and are likely capable of fatally attacking a person, but they, for the most part, are shy, are easily frightened and will avoid humans. Injuries caused by bears are rare, but are often widely reported.[31] The danger that bears pose is often vastly exaggerated, in part by the human imagination. However, when a mother feels her cubs are threatened, she will behave ferociously. It is recommended to give all bears a wide berth because they are behaviorally unpredictable.
Where bears raid crops or attack livestock, they may come into conflict with humans.[32][33] These problems may be the work of only a few bears, but they create a climate of conflict, as farmers and ranchers may perceive all losses as due to bears and advocate the preventive removal of all bears.[33] Mitigation methods may be used to reduce bear damage to crops, and reduce local antipathy towards bears.[32]
Bear danger area closure sign of a type used in Denali National Park in Alaska
Wojtek the bear with a Polish soldier: During the Battle of Monte Cassino, Wojtek helped move ammunition.
Laws have been passed in many areas of the world to protect bears from habitat destruction. Public perception of bears is often very positive, as people identify with bears due to their omnivorous diets, ability to stand on two legs, and symbolic importance,[34] and support for bear protection is widespread, at least in more affluent societies.[35] In more rural and poorer regions, attitudes may be more shaped by the dangers posed by bears and the economic costs they cause to farmers and ranchers.[33] Some populated areas with bear populations have also outlawed the feeding of bears, including allowing them access to garbage or other food waste. Bears in captivity have been trained to dance, box, or ride bicycles; however, this use of the animals became controversial in the late 20th century. Bears were kept for baiting in Europe at least since the 16th century.
Bear hunt
Some cultures use bears for food and folk medicine. Their meat is dark and stringy, like a tough cut of beef. In Cantonese cuisine, bear paws are considered a delicacy. The peoples of China, Japan, and Korea use bears' body parts and secretions (notably their gallbladders and bile) as part of traditional Chinese medicine. More than 12,000 bile bears are thought to be kept on farms, for their bile, in China, Vietnam, and South Korea.[36] Bear meat must be cooked thoroughly, as it can be infected with Trichinella spiralis, which can cause trichinosis.[37][38][39]
Culture
Names
The female first name "Ursula", originally derived from a Christian saint's name and common in English- and German-speaking countries, means "little she-bear" (diminutive of Latin ursa). In Switzerland, the male first name "Urs" is especially popular, while the name of the canton and city of Bern is derived from Bär, German for bear.
In Scandinavia, the male personal names Björn (Sweden, Iceland) and Bjørn (Norway, Denmark), meaning "bear", are relatively common. In Finland, the male personal name Otso is an old poetic name for bear, similar to Kontio.
In Russian and other Slavic languages, the word for bear, medved (медведь), and variants or derivatives such as Medvedev, are common surnames.
The Irish family name "McMahon" means "Son of Bear" in Irish.
In East European Jewish communities, the name Ber (בער)—Yiddish cognate of "Bear"—has been attested as a common male first name, at least since the 18th century, and was, among others, the name of several prominent rabbis. The Yiddish Ber is still in use among Orthodox Jewish communities in Israel, the US, and other countries. With the transition from Yiddish to Hebrew under the influence of zionism, the Hebrew word for "bear", dov (דב), was taken up in contemporary Israel and is at present among the commonly used male first names in that country.
"Ten Bears" (Paruasemana) was the name of a well-known 19th century chieftain among the Comanche. Also among other Native American tribes, bear-related names are attested.
Myth and legend
For bears in mythology, see Jean de l'Ours, Arcturus, Ursa Major, and Berserker.
There is evidence of prehistoric bear worship. Anthropologists such as Joseph Campbell have regarded this as a common feature in most of the fishing and hunting-tribes. The prehistoric Finns, along with most Siberian peoples, considered the bear as the spirit of one's forefathers. This is why the bear (karhu) was a greatly respected animal, with several euphemistic names (such as otso, mesikämmen and kontio). The bear is the national animal of Finland.
This kind of attitude is reflected in the traditional Russian fairy tale "Morozko", whose arrogant protagonist Ivan tries to kill a mother bear and her cubs—and is punished and humbled by having his own head turned magically into a bear's head and being subsequently shunned by human society.
"The Brown Bear of Norway" is a Scottish fairy tale telling the adventures of a girl who married a prince magically turned into a bear, and who managed to get him back into a human form by the force of her love and after many trials and difficulties. In the 1970s, this story was adapted into the East German fantasy film The Singing Ringing Tree and broadcast on British television.
Evidence of bear worship has been found in early Chinese and Ainu cultures, as well (see Iomante). Korean people in their mythology identify the bear as their ancestor and symbolic animal. According to the Korean legend, a god imposed a difficult test on a she-bear; when she passed it, the god turned her into a woman and married her.
Legends of saints taming bears are common in the Alpine zone. In the arms of the bishopric of Freising, the bear is the dangerous totem animal tamed by St. Corbinian and made to carry his civilised baggage over the mountains. A bear also features prominently in the legend of St. Romedius, who is also said to have tamed one of these animals and had the same bear carry him from his hermitage in the mountains to the city of Trento.
Similar stories are told of Saint Gall and Saint Columbanus.
This recurrent motif was used by the Church as a symbol of the victory of Christianity over paganism.[40] In the Norse settlements of northern England during the 10th century, a type of "hogback" grave cover of a long narrow block of stone, with a shaped apex like the roof beam of a long house, is carved with a muzzled, thus Christianised, bear clasping each gable end. Though the best collection of these is in the church at Brompton, North Yorkshire,[41] their distribution ranges across northern England and southern Scotland, with a scattered few in the north Midlands and single survivals in Wales, Cornwall, and Ireland; a late group is found in the Orkney Islands.
Bears are a popular feature of many children's stories, including Goldilocks and "The Story of the Three Bears", the Berenstein Bears, and Winnie the Pooh.
"En uheldig bjørnejakt" (An Unfortunate Bear Hunt) by Theodor Kittelsen.
Onikuma from Ehon Hyaku Monogatari
According to his hagiography, a bear killed Saint Corbinian's pack horse on the way to Rome, so the saint commanded it to carry his load. Once he arrived in Rome, however, he let the bear go.
The saddled "bear of St. Corbinian" the emblem of Freising, here incorporated in the arms of Pope Benedict XVI
Coat of Arms of the Abbey of Saint Gall
"The Three Bears", Arthur Rackham's illustration to English Fairy Tales, by
Symbolic use
For more details on this topic, see Bear in heraldry.
The British Lion, the Persian Cat and the Russian Bear (see The Great Game)
The Russian bear is a common national personification for Russia (as well as the Soviet Union). The brown bear is also Finland's national animal.
The flag of California
In the United States, the black bear is the state animal of Louisiana, New Mexico, and West Virginia; the grizzly bear is the state animal of both Montana and California. Bears also appear in the state seals of California and Missouri.
In the UK, the bear and staff has long featured on the heraldic arms of Warwickshire county.[42]
Bears appear in the canting arms of Bern and Berlin.
Also, "bear", "bruin", or specific types of bears are popular nicknames or mascots, for example, for sports teams (Chicago Bears, California Golden Bears, UCLA Bruins, Boston Bruins); and a bear cub called Misha was mascot of the 1980 Summer Olympics in Moscow, USSR.
Smokey Bear
Smokey Bear has become a part of American culture since his introduction in 1944. Known to almost all Americans, he and his message, "Only you can prevent forest fires" (updated in 2001 to "Only you can prevent wildfires"), have been a symbol of preserving woodlands.[43] Smokey wears a hat similar to one worn by U.S. Forest Service rangers; state police officers in some states wear a similar style, giving rise to the CB slang "bear" or "Smokey" for the highway patrol.
The name Beowulf has been hypothesized to mean "bee-wolf", a kenning for "bear".[44]
Figures of speech
The physical attributes and behaviours of bears are commonly used in figures of speech in English.
In the stock market, a bear market is a period of declining prices. Pessimistic forecasting or negative activity is said to be bearish (due to the stereotypical posture of bears looking downwards), and one who expresses bearish sentiment is a bear. Its opposite is a bull market, and bullish sentiment from bulls.
In gay slang, the term "bear" refers to male individuals who possess physical attributes much like a bear, such as a heavy build, abundant body hair, and commonly facial hair.
A bear hug is typically a tight hug that involves wrapping one's arms around another person, often leaving that person's arms immobile. It was used in the Ronald Reagan political ad "Bear in the woods".
Bear tracking – in the old Western states of the U.S. and, to this day, in the former Dakota Territory, the expression "you ain't just a bear trackin'" is used to mean "you ain't lying" or "that's for sure". This expression evolved as an outgrowth of the experience pioneer hunters and mountainmen had when tracking bear. Bears often lay down false tracks and are notorious for doubling back on anything tracking them. If you are not following bear tracks, you are not following false trails or leads in your thoughts, words or deeds.
In Korean culture a person is referred to as being "like a bear" when they are stubborn or not sensitive to what is happening around their surroundings. Used as a phrase to call a person "stubborn bear".
The Bible compares King David's "bitter warriors", who fight with such fury that they could overcome many times their number of opponents, with "a bear robbed of her whelps in the field" (2 Samuel 17:8 s:Bible (King James)/2 Samuel#Chapter 17). The phrase "a bereaved bear" (דב שכול), derived from this Biblical source, is still used in the literary Hebrew of contemporary Israel.
Teddy bears
Main article: Teddy bear
Around the world, many children have stuffed toys in the form of bears.
Organizations regarding bears
There are at least two authoritative organizations for seeking scientific information on bear species of the world. These focus on, for example, the species' natural history, management, and conservation. The International Association for Bear Research & Management is also known as the International Bear Association (IBA). The Bear Specialist Group of the Species Survival Commission is part of the International Union for the Conservation of Nature.
See also
Bear hunting
List of fatal bear attacks in North America
List of fictional bears
List of non-fictional bears
Ursari
Ursa major
Ursa minor
References
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^ abcdKrause, J.; Unger, T.; Noçon, A.; Malaspinas, A.; Kolokotronis, S.; Stiller, M.; Soibelzon, L.; Spriggs, H.; Dear, P. H.; Briggs, A. W.; Bray, S. C. E.; O'Brien, S. J.; Rabeder, G.; Matheus, P.; Cooper, A.; Slatkin, M.; Pääbo, S.; Hofreiter, M. (2008-07-28). "Mitochondrial genomes reveal an explosive radiation of extinct and extant bears near the Miocene-Pliocene boundary". BMC Evolutionary Biology8 (220): 220. doi:10.1186/1471-2148-8-220. PMC 2518930. PMID 18662376.
^Soibelzon, L. H.; Tonni, E. P.; Bond, M. (October 2005). "The fossil record of South American short-faced bears (Ursidae, Tremarctinae)". Journal of South American Earth Sciences20 (1–2): 105–113. doi:10.1016/j.jsames.2005.07.005.
^Qiu Zhanxiang (2003). "Dispersals of Neogene Carnivorans between Asia and North America" (PDF). Bulletin American Museum of Natural History279 (279): 18–31. doi:10.1206/0003-0090(2003)279<0018:C>2.0.CO;2. Archived from the original on 2009-11-20.
^Kurtén, B., 1995. The cave bear story: life and death of a vanished animal, Columbia University Press, ISBN 0-231-10361-1
^Lindburg, Donald G. (2004). Giant Pandas: Biology and Conservation, pp. 7–9. University of California Press, ISBN 0-520-23867-2
^Olaf R. P. Bininda-Emonds. "Phylogenetic Position of the Giant Panda". In Lindburg, Donald G. (2004) Giant Pandas: Biology and Conservation, pp. 11–35. University of California Press, ISBN 0-520-23867-2
^Flynn, J. J.; Nedbal, M. A.; Dragoo, J. W.; Honeycutt, R. L. (2000). "Whence the Red Panda?". Molecular Phylogenetics and Evolution17 (2): 190–199. doi:10.1006/mpev.2000.0819. PMID 11083933. Retrieved 2009-09-23.
^Flynn JJ, Finarelli JA, Zehr S, Hsu J, Nedbal MA (2005). "Molecular phylogeny of the carnivora (mammalia): assessing the impact of increased sampling on resolving enigmatic relationships". Systematic Biology54 (2): 317–337. doi:10.1080/10635150590923326. PMID 16012099.
^Flynn, J. J.; Nedbal, M. A. (1998). "Abstract". Molecular Phylogenetics and Evolution9 (3): 414–426. doi:10.1006/mpev.1998.0504. PMID 9667990. Archived from the original on 2010-11-18.. Retrieved 2009-10-08.
^"The Brown Bear: Father of the Polar Bear?, Alaska Science Forum". Gi.alaska.edu. 1996-12-05. Archived from the original on 2010-01-17. Retrieved 2010-10-16.
^Wood, Gerald (1983). The Guinness Book of Animal Facts and Feats. ISBN 978-0-85112-235-9.
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^ abCarnivores of the World by Dr. Luke Hunter. Princeton University Press (2011), ISBN 978-0-691-15228-8
^ abNovak, R. M. 1999. Walker's Mammals of the World. 6th edition. Johns Hopkins University Press, Baltimore. ISBN 0-8018-5789-9
^Bunnell, Fred (1984). In Macdonald, D. The Encyclopedia of Mammals. New York: Facts on File. p. 87. ISBN 978-0-87196-871-5.
^Mattson, David J. (1998). "Diet and Morphology of Extant and Recently Extinct Northern Bears". Ursus, A Selection of Papers from the Tenth International Conference on Bear Research and Management, Fairbanks, Alaska, July 1995, and Mora, Sweden, September 199510: 479–496. JSTOR 3873160.
^Ryan, Christopher; Pack, James C.; Igo, William K.; Billings, Anthony (2007). "Influence of mast production on black bear non-hunting mortalities in West Virginia". Ursus18 (1): 46–53. doi:10.2192/1537-6176(2007)18[46:IOMPOB]2.0.CO;2.
^Matheus, Paul E. (1995). "Diet and Co-ecology of Pleistocene Short-Faced Bears and Brown Bears in Eastern Beringia". Quaternary Research44 (3): 447–453. doi:10.1006/qres.1995.1090.
^Joshi, Anup; Garshelis, David L.; Smith, James L. D. (1997). "Seasonal and Habitat-Related Diets of Sloth Bears in Nepal". Journal of Mammalology1978 (2): 584–597. doi:10.2307/1382910.
^ abHwang, Mei-Hsiu (2002). "Diets of Asiatic black bears in Taiwan, with Methodological and Geographical Comparisons". Ursus13: 111–125.
^Zager, Peter; Beecham, John (2006). "The role of American black bears and brown bears as predators on ungulates in North America". Ursus17 (2): 95–108. doi:10.2192/1537-6176(2006)17[95:TROABB]2.0.CO;2.
^Clark, Douglas (2003). "Polar Bear–Human Interactions in Canadian National Parks, 1986–2000". Ursus14 (1): 65–71.
^ abFredriksson, Gabriella (2005). "Human–sun bear conflicts in East Kalimantan, Indonesian Borneo". Ursus16 (1): 130–137. doi:10.2192/1537-6176(2005)016[0130:HBCIEK]2.0.CO;2.
^Kellert, Stephen (1994). "Public Attitudes toward Bears and Their Conservation". Bears: Their Biology and Management9 (1): 43–50. doi:10.2307/3872683. JSTOR 3872683.
^Andersone, Žanete; Ozolinš, Jānis (2004). "Public perception of large carnivores in Latvia". Ursus15 (2): 181–187. doi:10.2192/1537-6176(2004)015<0181:PPOLCI>2.0.CO;2.
^Black, Richard (2007-06-11). "BBC Test kit targets cruel bear trade". BBC News. Archived from the original on 2010-11-18. Retrieved 2010-01-01.
^"Trichinellosis Associated with Bear Meat". Archived from the original on 30 September 2006. Retrieved 2006-10-04.
^"Bear meat poisoning in Siberia". BBC News. 1997-12-21. Retrieved 2006-10-04.
^"Finnish Food Safety Authority: Bear meat must be inspected before serving in restaurants". Retrieved 2006-10-04.
^Pastoreau, Michel (2007) L'ours. Historie d'un roi déchu, Seuil ISBN 2-02-021542-X.
^Noted and illustrated in Hall, Richard (1995) Viking Age Archaeology, p. 43 and fig. 22, ISBN 0-7478-0063-4.
^Warwickshire County Council, Civic heraldry
^"Forest Fire Prevention – Smokey Bear (1944–Present)". Ad Council. 1944-08-09. Archived from the original on 2010-11-18. Retrieved 2010-10-16.
^Sweet, Henry (1884) Anglo-Saxon Reader in Prose and Verse. The Clarendon Press, p. 202.
Further reading
Bears of the World, Terry Domico, photographs by Terry Domico and Mark Newman, Facts on File, Inc., 1988, hardcover, ISBN 978-0-8160-1536-8.
The Bear by William Faulkner.
Brunner, Bernd: Bears: A Brief History. New Haven and London: Yale University Press, 2007.
External links
Find more about Bears at Wikipedia's sister projects
Definitions and translations from Wiktionary
Media from Commons
Learning resources from Wikiversity
News stories from Wikinews
Quotations from Wikiquote
Source texts from Wikisource
Textbooks from Wikibooks
Species directories Ursidae from Wikispecies
The Bears Project – Information, reports and images of European brown bears and other living species
Western Wildlife Outreach – Information on the history, biology, and conservation of North American Grizzly Bears and Black Bears
v
t
e
Extant Carnivora species
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Infraclass: Eutheria
Superorder: Laurasiatheria
Suborder Feliformia
Nandiniidae
Nandinia
African palm civet (N. binotata)
Herpestidae (Mongooses)
Atilax
Marsh mongoose (A. paludinosus)
Bdeogale
Bushy-tailed mongoose (B. crassicauda)
Jackson's mongoose (B. jacksoni)
Black-footed mongoose (B. nigripes)
Crossarchus
Alexander's kusimanse (C. alexandri)
Angolan kusimanse (C. ansorgei)
Common kusimanse (C. obscurus)
Flat-headed kusimanse (C. platycephalus)
Cynictis
Yellow mongoose (C. penicillata)
Dologale
Pousargues's mongoose (D. dybowskii)
Galerella
Angolan slender mongoose (G. flavescens)
Somalian slender mongoose (G. ochracea)
Cape gray mongoose (G. pulverulenta)
Slender mongoose (G. sanguinea)
Helogale
Ethiopian dwarf mongoose (H. hirtula)
Common dwarf mongoose (H. parvula)
Herpestes
Short-tailed mongoose (H. brachyurus)
Indian gray mongoose (H. edwardsii)
Indian brown mongoose (H. fuscus)
Egyptian mongoose (H. ichneumon)
Small Asian mongoose (H. javanicus)
Long-nosed mongoose (H. naso)
Collared mongoose (H. semitorquatus)
Ruddy mongoose (H. smithii)
Crab-eating mongoose (H. urva)
Stripe-necked mongoose (H. vitticollis)
Ichneumia
White-tailed mongoose (I. albicauda)
Liberiictus
Liberian mongoose (L. kuhni)
Mungos
Gambian mongoose (M. gambianus)
Banded mongoose (M. mungo)
Paracynictis
Selous' mongoose (P. selousi)
Rhynchogale
Meller's mongoose (R. melleri)
Suricata
Meerkat (S. suricatta)
Hyaenidae (Hyenas)
Crocuta
Spotted hyena (C. crocuta)
Hyaena
Brown hyena (H. brunnea)
Striped hyena (H. hyaena)
Proteles
Aardwolf (P. cristatus)
Felidae
Large family listed below
Viverridae
Large family listed below
Eupleridae
Small family listed below
Family Felidae
Felinae
Acinonyx
Cheetah (A. jubatus)
Caracal
Caracal (C. caracal)
Felis
Chinese mountain cat (F. bieti)
Domestic cat (F. catus)
Jungle cat (F. chaus)
Pallas's cat (F. manul)
Sand cat (F. margarita)
Black-footed cat (F. nigripes)
Wildcat (F. silvestris)
Leopardus
Pantanal cat (L. braccatus)
Colocolo (L. colocolo)
Geoffroy's cat (L. geoffroyi)
Kodkod (L. guigna)
Andean mountain cat (L. jacobitus)
Pampas cat (L. pajeros)
Ocelot (L. pardalis)
Oncilla (L. tigrinus)
Margay (L. wiedii)
Leptailurus
Serval (L. serval)
Lynx
Canadian lynx (L. canadensis)
Eurasian lynx (L. lynx)
Iberian lynx (L. pardinus)
Bobcat (L. rufus)
Pardofelis
Marbled cat (P. marmorata)
Bay cat (P. badia)
Asian golden cat (P. temminckii)
Prionailurus
Leopard cat (P. bengalensis)
Iriomote cat (P. iriomotensis)
Flat-headed cat (P. planiceps)
Rusty-spotted cat (P. rubiginosus)
Fishing cat (P. viverrinus)
Profelis
African golden cat (P. aurata)
Puma
Cougar (P. concolor)
Jaguarundi (P. yagouaroundi)
Pantherinae
Panthera
Lion (P. leo)
Jaguar (P. onca)
Leopard (P. pardus)
Tiger (P. tigris)
Uncia
Snow leopard (U. uncia)
Neofelis
Clouded leopard (N. nebulosa)
Sunda clouded leopard (N. diardi)
Family Viverridae (includes Civets)
Paradoxurinae
Arctictis
Binturong (A. binturong)
Arctogalidia
Small-toothed palm civet (A. trivirgata)
Macrogalidia
Sulawesi palm civet (M. musschenbroekii)
Paguma
Masked palm civet (P. larvata)
Paradoxurus
Golden wet-zone palm civet (P. aureus)
Asian palm civet (P. hermaphroditus)
Jerdon's palm civet (P. jerdoni)
Golden palm civet (P. zeylonensis)
Hemigalinae
Chrotogale
Owston's palm civet (C. owstoni)
Cynogale
Otter civet (C. bennettii)
Diplogale
Hose's palm civet (D. hosei)
Hemigalus
Banded palm civet (H. derbyanus)
Prionodontinae (Asiatic linsangs)
Prionodon
Banded linsang (P. linsang)
Spotted linsang (P. pardicolor)
Viverrinae
Civettictis
African civet (C. civetta)
Genetta (Genets)
Abyssinian genet (G. abyssinica)
Angolan genet (G. angolensis)
Bourlon's genet (G. bourloni)
Crested servaline genet (G. cristata)
Common genet (G. genetta)
Johnston's genet (G. johnstoni)
Rusty-spotted genet (G. maculata)
Pardine genet (G. pardina)
Aquatic genet (G. piscivora)
King genet (G. poensis)
Servaline genet (G. servalina)
Haussa genet (G. thierryi)
Cape genet (G. tigrina)
Giant gorest genet (G. victoriae)
Poiana
African linsang (P. richardsonii)
Leighton's linsang (P. leightoni)
Viverra
Malabar large-spotted civet (V. civettina)
Large-spotted civet (V. megaspila)
Malayan civet (V. tangalunga)
Large Indian civet (V. zibetha)
Viverricula
Small Indian civet (V. indica)
Family Eupleridae
Euplerinae
Cryptoprocta
Fossa (C. ferox)
Eupleres
Falanouc (E. goudotii)
Fossa
Malagasy civet (F. fossana)
Galidiinae
Galidia
Ring-tailed mongoose (G. elegans)
Galidictis
Broad-striped Malagasy mongoose (G. fasciata)
Grandidier's mongoose (G. grandidieri)
Mungotictis
Narrow-striped mongoose (M. decemlineata)
Salanoia
Brown-tailed mongoose (S. concolor)
Durrell's vontsira (S. durrelli)
Suborder Caniformia (cont. below)
Ursidae (Bears)
Ailuropoda
Giant panda (A. melanoleuca)
Helarctos
Sun bear (H. malayanus)
Melursus
Sloth bear (M. ursinus)
Tremarctos
Spectacled bear (T. ornatus)
Ursus
American black bear (U. americanus)
Brown bear (U. arctos)
Polar bear (U. maritimus)
Asian black bear (U. thibetanus)
Mephitidae (Skunks)
Conepatus
(Hog-nosed
skunks)
Molina's hog-nosed skunk (C. chinga)
Humboldt's hog-nosed skunk (C. humboldtii)
American hog-nosed skunk (C. leuconotus)
Striped hog-nosed skunk (C. semistriatus)
Mephitis
Hooded skunk (M. macroura)
Striped skunk (M. mephitis)
Mydaus
Sunda stink badger (M. javanensis)
Palawan stink badger (M. marchei)
Spilogale (Spotted skunks)
Southern spotted skunk (S. angustifrons)
Western spotted skunk (S. gracilis)
Eastern spotted skunk (S. putorius)
Pygmy spotted skunk (S. pygmaea)
Procyonidae
Bassaricyon (Olingos)
Allen's olingo (B. alleni)
Beddard's olingo (B. beddardi)
Bushy-tailed olingo (B. gabbii)
Harris's olingo (B. lasius)
Chiriqui olingo (B. pauli)
Bassariscus
Ring-tailed cat (B. astutus)
Cacomistle (B. sumichrasti)
Nasua (Coatis inclusive)
White-nosed coati (N. narica)
South American coati (N. nasua)
Nasuella (Coatis inclusive)
Western mountain coati (N. olivacea)
Eastern mountain coati (N. meridensis)
Potos
Kinkajou (P. flavus)
Procyon
Crab-eating raccoon (P. cancrivorus)
Raccoon (P. lotor)
Cozumel raccoon (P. pygmaeus)
Ailuridae
Ailurus
Red panda (A. fulgens)
Suborder Caniformia (cont. above)
Otariidae
(Eared seals) (includes fur seals
and sea lions)
(Pinniped inclusive)
Arctocephalus
South American fur seal (A. australis)
Australasian fur seal (A. forsteri)
Galápagos fur seal (A. galapagoensis)
Antarctic fur seal (A. gazella)
Juan Fernández fur seal (A. philippii)
Brown fur seal (A. pusillus)
Guadalupe fur seal (A. townsendi)
Subantarctic fur seal (A. tropicalis)
Callorhinus
Northern fur seal (C. ursinus)
Eumetopias
Steller sea lion (E. jubatus)
Neophoca
Australian sea lion (N. cinerea)
Otaria
South American sea lion (O. flavescens)
Phocarctos
New Zealand sea lion (P. hookeri)
Zalophus
California sea lion (Z. californianus)
Galápagos sea lion (Z. wollebaeki)
Odobenidae (Pinniped inclusive)
Odobenus
Walrus (O. rosmarus)
Phocidae
(Earless seals)
(Pinniped inclusive)
Cystophora
Hooded seal (C. cristata)
Erignathus
Bearded seal (E. barbatus)
Halichoerus
Gray seal (H. grypus)
Histriophoca
Ribbon seal (H. fasciata)
Hydrurga
Leopard seal (H. leptonyx)
Leptonychotes
Weddell seal (L. weddellii)
Lobodon
Crabeater seal (L. carcinophagus)
Mirounga (Elephant seals)
Northern elephant seal (M. angustirostris)
Southern elephant seal (M. leonina)
Monachus
Mediterranean monk seal (M. monachus)
Hawaiian monk seal (M. schauinslandi)
Ommatophoca
Ross seal (O. rossi)
Pagophilus
Harp seal (P. groenlandicus)
Phoca
Spotted seal (P. largha)
Harbor seal (P. vitulina)
Pusa
Caspian seal (P. caspica)
Ringed seal (P. hispida)
Baikal seal (P. sibirica)
Canidae
Large family listed below
Mustelidae
Large family listed below
Family Canidae
Atelocynus
Short-eared dog (A. microtis)
Canis
Side-striped jackal (C. adustus)
Golden jackal (C. aureus)
Coyote (C. latrans)
Gray wolf (C. lupus)
Dog (C. lupus familiaris)
Black-backed jackal (C. mesomelas)
Red wolf (C. rufus)
Ethiopian wolf (C. simensis)
Cerdocyon
Crab-eating fox (C. thous)
Chrysocyon
Maned wolf (C. brachyurus)
Cuon
Dhole (C. alpinus)
Lycalopex
Culpeo (L. culpaeus)
Darwin's fox (L. fulvipes)
South American gray fox (L. griseus)
Pampas fox (L. gymnocercus)
Sechuran fox (L. sechurae)
Hoary fox (L. vetulus)
Lycaon
African wild dog (L. pictus)
Nyctereutes
Raccoon dog (N. procyonoides)
Otocyon
Bat-eared fox (O. megalotis)
Speothos
Bush dog (S. venaticus)
Urocyon
Gray fox (U. cinereoargenteus)
Island fox (U. littoralis)
Vulpes
Bengal fox (V. bengalensis)
Blanford's fox (V. cana)
Cape fox (V. chama)
Corsac fox (V. corsac)
Tibetan sand fox (V. ferrilata)
Arctic fox (V. lagopus)
Kit fox (V. macrotis)
Pale fox (V. pallida)
Rüppell's fox (V. rueppelli)
Swift fox (V. velox)
Red fox (V. vulpes)
Fennec fox (V. zerda)
Family Mustelidae
Lutrinae (Otters)
Aonyx
African clawless otter (A. capensis)
Oriental small-clawed otter (A. cinerea)
Enhydra
Sea otter (E. lutris)
Hydrictis
Spotted-necked otter (H. maculicollis)
Lontra
North American river otter (L. canadensis)
Marine otter (L. felina)
Neotropical otter (L. longicaudis)
Southern river otter (L. provocax)
Lutra
European otter (L. lutra)
Hairy-nosed otter (L. sumatrana)
Lutrogale
Smooth-coated otter (L. perspicillata)
Pteronura
Giant otter (P. brasiliensis)
Mustelinae (including badgers)
Arctonyx
Hog badger (A. collaris)
Eira
Tayra (E. barbara)
Galictis
Lesser grison (G. cuja)
Greater grison (G. vittata)
Gulo
Wolverine (G. gulo)
Ictonyx
Saharan striped polecat (I. libyca)
Striped polecat (I. striatus)
Lyncodon
Patagonian weasel (L. patagonicus)
Martes (Martens)
American marten (M. americana)
Yellow-throated marten (M. flavigula)
Beech marten (M. foina)
Nilgiri marten (M. gwatkinsii)
European pine marten (M. martes)
Japanese marten (M. melampus)
Fisher (M. pennanti)
Sable (M. zibellina)
Meles
Japanese badger (M. anakuma)
Asian badger (M. leucurus)
European badger (M. meles)
Mellivora
Honey badger (M. capensis)
Melogale (Ferret-badgers)
Bornean ferret-badger (M. everetti)
Chinese ferret-badger (M. moschata)
Javan ferret-badger (M. orientalis)
Burmese ferret-badger (M. personata)
Mustela (Weasels)
Amazon weasel (M. africana)
Mountain weasel (M. altaica)
Stoat (M. erminea)
Steppe polecat (M. eversmannii)
Colombian weasel (M. felipei)
Long-tailed weasel (M. frenata)
Japanese weasel (M. itatsi)
Yellow-bellied weasel (M. kathiah)
European mink (M. lutreola)
Indonesian mountain weasel (M. lutreolina)
Black-footed ferret (M. nigripes)
Least weasel (M. nivalis)
Malayan weasel (M. nudipes)
European polecat (M. putorius)
Siberian weasel (M. sibirica)
Back-striped weasel (M. strigidorsa)
Egyptian weasel (M. subpalmata)
Neovison (Minks)
American mink (N. vison)
Poecilogale
African striped weasel (P. albinucha)
Taxidea
American badger (T. taxus)
Vormela
Marbled polecat (V. peregusna)
v
t
e
Heraldic creatures
Beasts
Bear
Boar
Bull/Ox
Dog
Camelopard (giraffe)
Hind/Stag (deer)
Kangaroo
Leopard
Lion
Tiger
Wolf
Birds
Cock
Dove
Eagle
Martlet
Pelican
Rook
Legendary
creatures
Basilisk
Biscione
Chollima
Cockatrice
Dragon
Enfield
Garuda
Griffin/Keythong
Harpy
Lindworm
Manticore
Mermaid
Pegasus
Phoenix
Salamander
Sea-horse
Sea-lion
Turul
Unicorn
Wyvern
Fish
Dolphin
Ged
Hippocampus
Lucy (esox)
Scallop
Others
Bat
Bee
Crapaudy (toad)
Emmet (ant)
Serpent
UpToDate Contents
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Influence of different polymers on crystallization tendency and dissolution behavior of cilnidipine in solid dispersions.
Chen C1, Xie X, Li Y, Zhou C, Song Y, Yan Z, Yang X.Author information 1Laboratory of Pharmaceutics, Jiangsu Key Laboratory for Molecular and Medical Biotechnology, College of Life Sciences, Nanjing Normal University , Nanjing , PR China and.AbstractAbstract Context: Cilnidipine (CN) is a novel dihydropyridine calcium antagonist that is practically insoluble in aqueous media and exhibits a low oral bioavailability or limited clinical efficacy. Objective: This study investigated the effects of three commercial and chemically diverse polymers - PVP, PVP/VA and Soluplus - on crystallization tendency and in vitro dissolution profiles of CN in order to determine an optimum carrier for composing the preferred solid dispersion (SD) of CN. Methods: All these co-evaporated systems were characterized up to 3 months by thermoanalytical (DSC), crystallographic (POM, PXRD), microscopic (SEM) and spectroscopic (FTIR) techniques. Results: The results showed that the polymers could be sorted by their effects of inhibiting CN crystallization in the ascending order: Soluplus, PVP/VA, PVP. The sequence was in accordance with that of the strength of drug-polymer hydrogen bonds revealed by FTIR spectra. It could be ascribed to relative hydrogen-bonding acceptor strengths of N-vinylpyrrolidone moiety in the polymer molecules. On the other hand, all the SDs showed enhanced dissolution profiles compared to pure CN alone. On their effects of enhancing CN dissolution, the polymers could be sorted in the descending order: Soluplus, PVP, PVP/VA. Conclusions: It implied that the dissolution behavior of CN could bear a close relationship to both hydration capacity and hydrogen-bonding interaction tendency of moieties of the polymers. It might suggest an optimal formulation for CN comprising both PVP and Soluplus.
Drug development and industrial pharmacy.Drug Dev Ind Pharm.2014 Apr;40(4):441-51. doi: 10.3109/03639045.2013.767825. Epub 2013 Apr 24.
Abstract Context: Cilnidipine (CN) is a novel dihydropyridine calcium antagonist that is practically insoluble in aqueous media and exhibits a low oral bioavailability or limited clinical efficacy. Objective: This study investigated the effects of three commercial and chemically diverse polymers - P
Early complications of surgery in operative treatment of ankle fractures in those over 60: A review of 186 cases.
Zaghloul A1, Haddad B2, Barksfield R3, Davis B3.Author information 1West Suffolk Hospital, Hardwick Lane Bury St. Edmunds, Suffolk IP33 2QZ, UK; Norfolk and Norwich University Hospital, Colney Lane, Norwich NR4 7UY, UK.2West Suffolk Hospital, Hardwick Lane Bury St. Edmunds, Suffolk IP33 2QZ, UK; University College London, Institute of Orthopaedic and Musculoskeletal Sciences, Royal National Orthopaedic Hospital Stanmore, Middlesex HA7 4LP, UK. Electronic address: behrooz.haddad@gmail.com.3Norfolk and Norwich University Hospital, Colney Lane, Norwich NR4 7UY, UK.AbstractINTRODUCTION: Ankle fractures are among the most common injuries of the lower extremity encountered by orthopaedic surgeons. With increasing population age and osteoporosis, the prevalence of these fractures is expected to increase. The aim of this study was to evaluate complications and the need for revision surgery after the surgical treatment of ankle fractures in patients over 60 years of age. We report the outcomes of 186 consecutive patients who underwent operative treatment for rotational ankle fractures in our institution from 2007 to 2010.
Injury.Injury.2014 Apr;45(4):780-3. doi: 10.1016/j.injury.2013.11.008. Epub 2013 Nov 23.
INTRODUCTION: Ankle fractures are among the most common injuries of the lower extremity encountered by orthopaedic surgeons. With increasing population age and osteoporosis, the prevalence of these fractures is expected to increase. The aim of this study was to evaluate complications and the need fo
APOBEC3 Multimerization Correlates with HIV-1 Packaging and Restriction Activity in Living Cells.
Li J1, Chen Y1, Li M2, Carpenter MA2, McDougle RM2, Luengas EM2, Macdonald PJ3, Harris RS2, Mueller JD4.Author information 1School of Physics and Astronomy, University of Minnesota, 116 Church Street Southeast, Minneapolis, MN 55455, USA; Institute for Molecular Virology, University of Minnesota, 515 Delaware Street Southeast, Minneapolis, MN 55455, USA.2Institute for Molecular Virology, University of Minnesota, 515 Delaware Street Southeast, Minneapolis, MN 55455, USA; Department of Biochemistry, Molecular Biology and Biophysics, 321 Church Street Southeast, Minneapolis, MN 55455, USA.3Department of Biomedical Engineering, University of Minnesota, 312 Church Street Southeast, Minneapolis, MN 55455, USA.4School of Physics and Astronomy, University of Minnesota, 116 Church Street Southeast, Minneapolis, MN 55455, USA; Institute for Molecular Virology, University of Minnesota, 515 Delaware Street Southeast, Minneapolis, MN 55455, USA; Department of Biomedical Engineering, University of Minnesota, 312 Church Street Southeast, Minneapolis, MN 55455, USA. Electronic address: Mueller@physics.umn.edu.AbstractAPOBEC3G belongs to a family of DNA cytosine deaminases that are involved in the restriction of a broad number of retroviruses including human immunodeficiency virus type 1 (HIV-1). Prior studies have identified two distinct mechanistic steps in Vif-deficient HIV-1 restriction: packaging into virions and deaminating viral cDNA. APOBEC3A, for example, although highly active, is not packaged and is therefore not restrictive. APOBEC3G, on the other hand, although having weaker enzymatic activity, is packaged into virions and is strongly restrictive. Although a number of studies have described the propensity for APOBEC3 oligomerization, its relevance to HIV-1 restriction remains unclear. Here, we address this problem by examining APOBEC3 oligomerization in living cells using molecular brightness analysis. We find that APOBEC3G forms high-order multimers as a function of protein concentration. In contrast, APOBEC3A, APOBEC3C and APOBEC2 are monomers at all tested concentrations. Among other members of the APOBEC3 family, we show that the multimerization propensities of APOBEC3B, APOBEC3D, APOBEC3F and APOBEC3H (haplotype II) bear more resemblance to APOBEC3G than to APOBEC3A/3C/2. Prior studies have shown that all of these multimerizing APOBEC3 proteins, but not the monomeric family members, have the capacity to package into HIV-1 particles and restrict viral infectivity. This correlation between oligomerization and restriction is further evidenced by two different APOBEC3G mutants, which are each compromised for multimerization, packaging and HIV-1 restriction. Overall, our results imply that multimerization of APOBEC3 proteins may be related to the packaging mechanism and ultimately to virus restriction.
Journal of molecular biology.J Mol Biol.2014 Mar 20;426(6):1296-307. doi: 10.1016/j.jmb.2013.12.014. Epub 2013 Dec 17.
APOBEC3G belongs to a family of DNA cytosine deaminases that are involved in the restriction of a broad number of retroviruses including human immunodeficiency virus type 1 (HIV-1). Prior studies have identified two distinct mechanistic steps in Vif-deficient HIV-1 restriction: packaging into virion
An efficient strategy for the synthesis of polysubstituted chromeno[4,3-b]pyrrolidine derivatives.
Tian L1, Xu GQ, Li YH, Liang YM, Xu PF.Author information 1State Key Laboratory of Applied Organic Chemistry, College of Chemistry and Chemical Engineering, Lanzhou University, Lanzhou 730000, P.R. China. liangym@lzu.edu.cn xupf@lzu.edu.cn.AbstractA powerful cascade reaction was developed for the synthesis of chromeno[4,3-b]pyrrolidines with high yields and excellent stereoselectivities. This efficient cascade reaction expeditiously established two fused rings which bear three contiguous stereogenic centers including one quaternary stereocenter in a single operation with low catalyst loading under mild conditions.
Chemical communications (Cambridge, England).Chem Commun (Camb).2014 Mar 7;50(19):2428-30. doi: 10.1039/c3cc49504c. Epub 2014 Jan 22.
A powerful cascade reaction was developed for the synthesis of chromeno[4,3-b]pyrrolidines with high yields and excellent stereoselectivities. This efficient cascade reaction expeditiously established two fused rings which bear three contiguous stereogenic centers including one quaternary stereocent
… While many host countries pay little to or some even receive money from the United States for hosting the United States forces, Japan is very generous to bear the cost to maintain USFJ up to 75% in spite of the fact that the Japanese Government suffers from huge budget deficit. …
