ウラボシ科
WordNet
- ferns: a large family that in some classification systems has been subdivided into several families (including Aspleniaceae and Blechnaceae and Davalliaceae and Dennstaedtiaceae and Dryopteridaceae and Oleandraceae and Pteridaceae) (同)family Polypodiaceae
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出典(authority):フリー百科事典『ウィキペディア（Wikipedia）』「2013/06/16 09:40:49」(JST)
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| Polypodiaceae |
|
| Pleopeltis polypodioides fronds on an oak limb |
| Scientific classification |
| Kingdom: |
Plantae |
| Division: |
Pteridophyta |
| Class: |
Polypodiopsida/Pteridopsida
(disputed) |
| Order: |
Polypodiales |
| (unranked): |
Eupolypods |
| (unranked): |
Eupolypods I |
| Family: |
Polypodiaceae |
| Subfamilies/Tribes/Clades |
|
Platycerioideae
Polypodioideae
- Drynarieae
- Selligueeae
- Lepisoreae
- Microsoreae
- Polypodieae
- Loxogrammeae
Grammitids
|
| Synonyms |
|
Drynariaceae
Grammitidaceae
Gymnogrammitidaceae
Loxogrammaceae
Platyceriaceae
Pleurisoriopsidaceae
|
Polypodiaceae is a family of polypod ferns, which includes more than 60 genera divided into several tribes and containing around 1,000 species. Nearly all are epiphytes, but some are terrestrial.[1]
|
Contents
- 1 Description
- 2 Classification
- 3 Phylogeny
- 4 Selected genera
- 5 See also
- 6 References
- 7 External links
|
Description[edit]
Stems of Polypodiaceae range from erect to long-creeping. The fronds are entire, pinnatifid, or variously forked or pinnate. The petioles lack stipules. The scaly rhizomes are generally creeping in nature. Polypodiaceae species are found in wet climates, most commonly in rain forests. In temperate zones, most species tend to be epiphytic or epipetric.[1]
Notable examples of ferns in this family include the resurrection fern (Pleopeltis polypodioides) and the golden serpent fern (Phlebodium aureum).[1]
Classification[edit]
Recent molecular phylogenic analysis has led to the division of the Polypodiaceae into five subfamiles, and to the inclusion of genera that have at various times been placed in other families, including the Drynariaceae, Grammitidaceae, Gymnogrammitidaceae, Loxogrammaceae, Platyceriaceae, and Pleurisoriopsidaceae.[2] In the list that follows, the taxa shown with the "(=)" prefix are considered to be synonymyms for the accepted subfamily name that they follow. However, this does not necessarily imply that the subfamily contains all of the synonym's previous genera.
- Subfamily Loxogrammoideae H.Schneid. 2011
- (=) tribe Loxogrammeae R.M.Tryon & A.F.Tryon 1982
- (=) family Loxogrammaceae Ching ex Pic.Serm. 1975
- Lacks sclerenchyma (supporting tissue) in plant body, except in the roots.[3]
- Genera: Dictymia, Loxogramme
- Subfamily Drynarioideae Crabbe, Jermy & Mickel 1975
- (=) tribe Drynarieae Subh.Chandra 1982
- (=) tribe Selligueeae (author?)[3]
- (=) family Drynariaceae Ching 1978
- Combines drynarioid and selligueoid ferns
- Genera: Aglaomorpha, Arthromeris, Christiopteris, Drynaria, Gymnogrammitis, Paraselliguea, Phymatopteris, Polypodiopteris, Selliguea
- Subfamily Platycerioideae B.K.Nayar 1970
- (=) family Platyceriaceae Ching 1978
- Fronds with stellate (radiating from central point) hairs.[3]
- Genera: Platycerium, Pyrrosia
- Subfamily Microsoroideae B.K.Nayar 1970 (as ‘Microsorioideae’)
- (=) tribe Microsoreae V.N.Tu 1981
- Genera: Dendroconche, Goniophlebium, Kaulinia, Kontumia, Lecanopteris, Lemmaphyllum, Lepisorus, Lepidomicrosorium, Leptochilus, Microsorum, Neocheiropteris, Neolepisorus, Paragramma, Phymatosorus, Podosorus, Thylacopteris, Tricholepidium
- Subfamily Polypodioideae B.K.Nayar 1970
- (=) tribe Polypodieae Hooker & Lindley ex Duby (sensu Mabberley 2008)
-
- Genera: Campyloneurum, Microgramma, Niphidium, Pecluma, Phlebodium, Pleopeltis, Pleurosoriopsis, Polypodium, Serpocaulon, Synammia
- Grammitid ferns (includes family Grammitidaceae)
-
- Genera: Acrosorus, Adenophorus, Calymmodon, Ceradenia, Chrysogrammitis, Cochlidium, Ctenopterella, Dasygrammitis, Enterosora, Grammitis, Lellingeria, Leucostrichum, Luisma, Melpomene, Micropolypodium, Oreogrammitis, Prosaptia (incl. Ctenopteris), Radiogrammitis, Scleroglossum (incl. Nematopteris), Terpsichore, Themelium, Tomophyllum, Xiphopterella, Xiphopteris, Zygophlebia
Phylogeny[edit]
The following phylogram shows a likely relationship between the Polypodiaceae subfamilies.[4]
| Polypodiaceae |
|
Loxogrammoideae
|
|
|
|
|
Drynarioideae
|
|
|
|
Platycerioideae
|
|
|
Microsoroideae
|
|
|
|
|
|
|
Polypodioideae
|
|
|
|
|
|
Selected genera[edit]
- Acrosorus
- Adenophorus
- Aglaomorpha - includes Holostachyum, Merinthosorus, Photinopteris, Pseudodrynaria
- Arthromeris
- Belvisia
- Calymmodon
- Campyloneurum - formerly included in Polypodium
- Caobangia
- Ceradenia
- Christiopteris
- Chrysogrammitis
- Cochlidium
- Colysis
- Ctenopteris
- Dicranoglossum
- Dictymia
- Drymotaenium
- Drynaria
- Enterosora
- Goniophlebium
- Grammitis
- Gymnogrammitis
- Kaulinia
- Kontumia
- Lecanopteris
- Lellingeria
- Lemmaphyllum
- Lepisorus
- Leptochilus - includes Platygyria
- Leucotrichum
- Lomaria
- Loxogramme - includes Anarthropteris
- Luisma
|
- Marginariopsis
- Melpomene
- Microgramma - includes Solanopteris, formerly included in Polypodium
- Microphlebodium
- Micropolypodium
- Microsorum
- Myrmecophila
- Neocheiropteris - includes Neolepisorus
- Neurodium - formerly included in Polypodium
- Niphidium
- Pecluma - formerly included in Polypodium
- Phlebodium - formerly included in Polypodium
- Phymatoides
- Phymatosorus - sometimes included in Polypodium
- Platycerium
- Pleopeltis - formerly included in Polypodium
- Pleopodium
- Pleurosoriopsis
- Podosorus
- Polypodiodes
- Polypodium
- Prosaptia
- Pseudocolysis
- Pyrrosia - includes Drymoglossum
- Scleroglossum
- Selliguea - includes Crypsinus, Polypodiopteris
- Serpocaulon
- Solanopteris
- Synammia
- Terpsichore
- Themelium
- Thylacopteris
- Zygophlebia
|
See also[edit]
- List of foliage plant diseases (Polypodiaceae)
References[edit]
- ^ a b c G. Panigrahi and S.N. Patnaik (1961), Cytology of Some Genera of Polypodiaceae in Eastern India, Nature 191, 1207 - 1208.
- ^ Christenhusz et al.: A linear sequence of extant families and genera of lycophytes and ferns Phytotaxa 19: 7-54. (18 Feb. 2011)
- ^ a b c D. J. Mabberley (2008). Mabberley's plant-book: a portable dictionary of plants, their classification and uses. Cambridge University Press. p. 690. ISBN 978-0-521-82071-4.
- ^ Schuettpelz (2008) Schuettpelz & Pryer "Fern pylogeny" in Biology and Evolution of Ferns and Lycophytes], ed. Tom A. Ranker and Christopher H. Haufler. Cambridge University Press 2008
External links[edit]
- Polypodiaceae (Polypody family)
English Journal
- Morphological innovation, ecological opportunity, and the radiation of a major vascular epiphyte lineage.
- Sundue MA1, Testo WL2, Ranker TA3.
- Evolution; international journal of organic evolution.Evolution.2015 Aug 8. doi: 10.1111/evo.12749. [Epub ahead of print]
- The emergence of angiosperm-dominated tropical forests in the Cretaceous led to major shifts in the composition of biodiversity on Earth. Among these was the rise to prominence of epiphytic plant lineages, which today comprise an estimated one-quarter of tropical vascular plant diversity. Among the
- PMID 26257202
- Higher clonal integration in the facultative epiphytic fern Selliguea griffithiana growing in the forest canopy compared with the forest understorey.
- Lu HZ1, Liu WY2, Yu FH3, Song L3, Xu XL3, Wu CS1, Zheng YL3, Li YP1, Gong HD3, Chen K1, Li S3, Chen X1, Qi JH3, Lu SG2.
- Annals of botany.Ann Bot.2015 Jul;116(1):113-22. doi: 10.1093/aob/mcv059. Epub 2015 Jun 6.
- BACKGROUND AND AIMS: The advantage of clonal integration (resource sharing between connected ramets of clonal plants) varies and a higher degree of integration is expected in more stressful and/or more heterogeneous habitats. Clonal facultative epiphytes occur in both forest canopies (epiphytic habi
- PMID 26050068
- Polypodium leucotomos: a potential new photoprotective agent.
- Bhatia N1.
- American journal of clinical dermatology.Am J Clin Dermatol.2015 Apr;16(2):73-9. doi: 10.1007/s40257-015-0113-0.
- As the understanding of the immune system pathways, cytokine balances, and cellular interactions continues to expand, so must the potential applications of therapies that can impact the process of diseases instead of just controlling their symptoms. In the case of Polypodium leucotomos extract, whic
- PMID 25666116
- Volatile organic compounds in the strongly fragrant fern genus Melpomene (Polypodiaceae).
- Kessler M1, Connor E, Lehnert M.
- Plant biology (Stuttgart, Germany).Plant Biol (Stuttg).2015 Mar;17(2):430-6. doi: 10.1111/plb.12252. Epub 2014 Nov 26.
- Volatile organic compounds (VOCs) are common among plants, both as attractants for pollinators and as defence against herbivores. While much studied among flowering plants, the prevalence and function of VOCs among ferns is little known. Using headspace sorption and gas chromatography, we analysed t
- PMID 25427549
Japanese Journal
- Evolution of Epiphytism in Ferns and Lycophytes with an Emphasis on Davalliaceae(<SPECIAL SERIES>INVITED ARTICLES BY SUCCESSIVE PRESIDENTS OF THE JAPANESE SOCIETY FOR PLANT SYSTEMATICS)
- KATO MASAHIRO,TSUTSUMI CHIE
- APG : Acta phytotaxonomica et geobotanica 64(3), 159-177, 2013-10-31
- … The family forms the largest, generally epiphytic monophyletic clade with Polypodiaceae including grammitids. … Epiphytic gametophytes are strap-shaped, ribbon-like or filamentous, branched, gemmiferous and long-lived in, e.g., grammitids, Hymenophyllaceae and vittarioids, while, like terrestrial gametophytes, they are heart-shaped and monocarpic in, e.g., Davalliaceae and Polypodiaceae. …
- NAID 110009661544
- WAS JEJU ISLAND A GLACIAL REFUGIUM FOR EAST ASIAN WARM-TEMPERATE PLANTS? INSIGHTS FROM THE HOMOSPOROUS FERN SELLIGUEA HASTATA (POLYPODIACEAE)
- Chung MY,Moon MO,Lopez-Pujol J,Maki M,Yamashiro T,Yukawa T,Sugiura N,Lee YI,Chung MG
- AMERICAN JOURNAL OF BOTANY 100(11), 2240-2249, 2013
- NAID 120005597795
- 428 Morphological analysis of extant Serpocaulon A. R. Sm. (Polypodiaceae J. Presl. & C. Presl.) spores from Colombia (NW South America) : a tool for the study of the Cenozoic Neotropical paleobotany
- Ramirez-Valencia Valentina,Sanin David,Pardo-Trujillo Andres
- 日本花粉学会会誌 58(特別), 194-195, 2012-08-23
- NAID 110009572591
- 日本・韓国・台湾のウラボシ科ミツデウラボシ属の新学名
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